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Siderolamprus enneagrammus COPE, 1861

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Higher TaxaDiploglossidae, Diploglossa, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Huaxteca Lesser Galliwasp
S: Celesto Huasteco 
SynonymSiderolamprus enneagrammus COPE 1861: 368
Diploglossus steindachneri COPE 1864 (fide SMITH & TAYLOR 1950)
Diploglossus chalybaeus COPE 1867: 321 (fide SMITH & TAYLOR 1950)
Celestus chalybaeus — COPE 1885: 170
Diploglossus steindachneri — GÜNTHER 1885: 34
Diploglossus steindachneri — BOULENGER 1885: 293
Celestus enneagrammus — COPE 1887
Celestus enneagrammus — SMITH & TAYLOR 1950: 195
Diploglossus enneagrammus — WERMUTH 1969
Celestus enneagrammus — GOOD 1987
Celestus enneagrammus — LINER 1994
Diploglossus enneagrammus — WERLER & CAMPBELL 2004
Celestus enneagrammus — MATA-SILVA et al. 2015
Siderolamprus enneagrammus — SCHOOLS & HEDGES 2021 
DistributionMexico (coastal areas of Oaxaca, Veracruz, Puebla, Chiapas)

Type locality: Jalapa, Veracruz.  
Reproductionovovivparous 
TypesHolotype: MCZ 2848
Holotype: USNM 6603 [chalybaeus]
Holotype: USNM 6342 [Diploglossus steindachneri] 
DiagnosisDiagnosis (genus Siderolamprus). Species of Siderolamprus have (1) claw sheath, absent, (2) contact between the nasal and rostral scales, present or absent, (3) scales in contact with the nasal scale, six, (4) postnasal scales, two, (5) position of the nostril in the nasal scale, posterior, (6) keels on dorsal body scales, present or absent, (7) digits per limb, five, (8) longest toe lamellae, 14–35, (9) dorsal scale rows, 65–85, (10) relative head width, 9.88–15.6, (11) relative rostral height, 44.7–63.9, (12) relative frontonasal length, 2.46–2.88, (13) relative interparietal distance, 0.561–1.14, (14) relative axilla-groin distance, 53.5–65.3. From Mesoamericus gen. nov., we distinguish Siderolamprus by the claw sheath (absent versus present in Mesoamericus) and relative rostral height (44.7–63.9 versus 65.0–67.0) (Schools & Hedges 2021). 
CommentType species: Siderolamprus enneagrammus COPE 1861: 368 is the type species of the genus Siderolamprus COPE 1861. Content: 14 species: Siderolamprus adercus, S. atitlanensis, S. bivittatus, S. cyanochloris, S. enneagrammus, S. hylaius, S. ingridae, S. laf, S. legnotus, S. montanus, S. orobius, S. owenii, S. rozellae, S. scansorius (fide Schools & Hedges 2021).

Habitat: Species of Siderolamprus are ground- or tree-adapted ecomorphs (Schools et al. 2022). 
EtymologyFrom the Greek “ennéa” meaning nine, “-gramma” meaning something written, and the Latin ending “-us” (masculine). Likely in reference to the nine longitudinal black lines along its body. 
References
  • Aguilar-López JL, Luría-Manzano R, Pineda E, Canseco-Márquez L 2021. Selva Zoque, Mexico: an important Mesoamerican tropical region for reptile species diversity and conservation. ZooKeys 1054: 127-153 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Campbell J A; Camarillo R J L 1994. A new lizard of the genus Diploglossus (Anguidae: Diploglossinae) from Mexico, with a review of the Mexican and northern Central American species. Herpetologica 50 (2): 193-209 - get paper here
  • Canseco-Marquez,L.; Gutierrez-Mayen,G. & Salazar-Arenas,J. 2000. New records and range extensions for amphibians and reptiles from Puebla, México. Herpetological Review 31 (4): 259-263 - get paper here
  • Casas-Andreu, G., F.R. Méndez-De la Cruz and X. Aguilar-Miguel. 2004. Anfibios y Reptiles; pp. 375–390, in A.J.M. García-Mendoza, J. Ordoñez and M. Briones-Salas (ed.). Biodiversidad de Oaxaca. Instituto de Biología, UNAM-Fondo Oaxaqueño para la Conservación de la Naturaleza-World Wildlife Fund, México, D. F.
  • CASITLLO-JUÁREZ, JORGE LUIS; VÍCTOR VÁSQUEZ-CRUZ & LAURA PAMELA TAVAL-VELÁZQUEZ. 2021. CELESTUS ENNEAGRAMMUS (Huaxteca Lesser Galliwasp). REPRODUCTION and MAXIMUM BODY SIZE. Herpetological Review 52 (3): 646–647. - get paper here
  • Cope, E.D. 1861. Descriptions of reptiles from tropical America and Asia. Proc. Acad. Nat. Sci. Philadelphia 1860: 368-374 - get paper here
  • Cope, E.D. 1867. Fifth contribution lo the herpetology of tropical America. Proc. Acad. nat. Sci. Philadelphia, 18 [1866]: 317-323 - get paper here
  • Good, D. A. 1987. An allozyme analysis of anguid subfamilial relationships (Lacertilia: Anguidae). Copeia 1987 (3): 696-701 - get paper here
  • Günther, A. C. L. G. 1885. Reptilia and Batrachia. Biologia Centrali-Américana. Taylor, & Francis, London, 326 pp. [published in parts from 1885-1902; reprint by the SSAR 1987] - get paper here
  • Johnson, Jerry D.; Vicente Mata-Silva, Elí García Padilla, and Larry David Wilson 2015. The Herpetofauna of Chiapas, Mexico: composition, distribution, and conservation. Mesoamerican Herpetology 2 (3): 272–329. - get paper here
  • Manuel, Feria-Ortiz and Serrano-Garcia Josè Rodrigo. 2016. Celestus enneagrammus (Huaxteca Lesser Galliwasp) Reproduction. Herpetological Review 47 (2): 295-296 - get paper here
  • Mata-Silva, Vicente, Jerry D. Johnson, Larry David Wilson and Elí García-Padilla. 2015. The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology 2 (1): 6–62 - get paper here
  • Ramírez González, Cynthia Grisell; Ángel Edgar Lara Brenis, Teresa de Jesús Mijangos Arrazola 2014. New Distributional Records of Amphibians and Reptiles from Northern Oaxaca, México. Check List 10 (3): 679-681 - get paper here
  • Savage, JM; Lips, KR; Ibáñez RD 2008. A new species of Celestus from west-central Panama, with consideration of the status of the genera of the Anguidae: Diploglossinae (Squamata). Revista de Biologia Tropical 56 (2): 845-859 - get paper here
  • Schools, M., Kasprowicz, A., & Hedges, S. B. 2022. Phylogenomic data resolve the historical biogeography and ecomorphs of Neotropical forest lizards (Squamata, Diploglossidae). Molecular Phylogenetics and Evolution, 107577 - get paper here
  • SCHOOLS, MOLLY & S. BLAIR HEDGES 2021. Phylogenetics, classification, and biogeography of the Neotropical forest lizards (Squamata, Diploglossidae). Zootaxa 4974 (2): 201–257 - get paper here
  • Smith, H.M. & Taylor,E.H. 1950. An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bull. US Natl. Mus. 199: 1-253 - get paper here
  • Stuart, L. C. 1948. The amphibians and reptiles of Alta Verapaz Guatamala. Miscellaneous Publications, Museum of Zoology, University of Michigan 69: 1-109 - get paper here
  • Torres-Hernández, LA, Ramírez-Bautista A, Cruz-Elizalde R, Hernández-Salinas U, Berriozabal-Islas C, DeSantis DL, Johnson JD, Rocha A, García-Padilla E, Mata-Silva V, Fucsko LA, and Wilson LD. 2021. The herpetofauna of Veracruz, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 15(2) [General Section]: 72–155 - get paper here
  • Werler, J.E. & Campbell, J.A. 2004. NEW LIZARD OF THE GENUS DIPLOGLOSSUS (ANGUIDAE: DIPLOGLOSSINAE) FROM THE TUXTLAN FAUNAL REGION, VERACRUZ, MEXICO. Southwestern Naturalist 49 (3):327–333 - get paper here
  • Woolrich-Piña, G. A., E. García-Padilla, D. L. DeSantis, J. D. Johnson, V. Mata-Silva, and L. D. Wilson 2017. The herpetofauna of Puebla, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 4(4): 791–884 - get paper here
 
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