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Trimeresurus truongsonensis ORLOV, RYABOV, THANH & CUC, 2004

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Quang Binh pitviper
G: Quang Binh-Bambusotter 
SynonymTrimeresurus truongsonensis ORLOV, RYABOV, THANH & CUC 2004
Viridovipera truongsonensis — DAWSON et al. 2008
Trimeresurus (Viridovipera) truongsonensis — DAVID et al. 2011
Viridovipera truongsonensis — WALLACH et al. 2014: 790
Viridovipera truongsonensis — GUO et al. 2018
Trimeresurus truongsonensis— RATHEE et al. 2022
Trimeresurus (Viridovipera) truongsonensis — MIRZA et al. 2023 
DistributionC Vietnam (May be more widely distributed in Quang Binh Province beyond the type locality and southwards to Quang Tri Prov.)

Type locality: Phong Nha, Ke Bang National Park, Quang Binh Province, 600 m, Vietnam.  
Reproductionovovivparous 
TypesHolotype: ZISP 22931 
DiagnosisDiagnosis: Slender, small Trimeresurus, up to 642 mm (SVL males is up to 521 mm, Lcd — 121 mm) with very hard, thin and almost round body. V — 166 – 175; Scd — 65 – 69 pairs; anal plate is entire. Scale rows oblique, rhomboid, strongly keeled, only first row of dorsal scales has hardly noticeable keels; Sq2 — 21, Sq3 — 15 rows. The head is small, triangular, clearly distinct from the neck, covered by small, strongly granulated shields on the upper surface backwards from the line between eyes, shields are relatively flat from snout to eyes. Supralabials — 9–10, Infralabials — 11–12; the first supralabial is clearly divided from large N; the rostral shield forms isosceles triangle with truncated top; SO — 1, very large; SubO — 1, thin and long; symphisial relatively large, triangular; anterior genial shields very large and wide, posterior genial shields small. Hemipenis relatively short (to 7th Scd) with large and small spines, forked. The dorsal coloration of the body and the head of males is greenish-blue, in females — light-brown. Across body and tail pass 66 — 70 red-brown bands; red-brown spots on the head form mosaic pattern. The belly is gray-bluish. (Orlov et al. 2004)


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CommentVenomous!

Characteristic features. Slender, small pitviper, with small, triangular head; 11 supralabials, with first supralabial clearly divided from large nasal; 13 infralabials, 167 ventrals, anal scale entire, 61 divided subcaudals, and 21 (dorsally keeled) scales across the midbody; about 72 dark bands on dorsum of neck, body and tail (from ZIEGLER et al. 2006).

Habitat: partly arboreal (Harrington et al. 2018). 
EtymologyNamed after the mountain range Truong Son (= Annamite mountains) where the species occurs. 
References
  • DAVID, PATRICK; GERNOT VOGEL & ALAIN DUBOIS 2011. On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacépède, 1804 (Reptilia: Squamata: Viperidae). Zootaxa 2992: 1–51 - get paper here
  • Dawson, Karen; Anita Malhotra, Roger S. Thorpe, Peng Guo, Mrinalini and Thomas Ziegler 2008. Mitochondrial DNA analysis reveals a new member of the Asian pitviper genus Viridovipera (Serpentes: Viperidae: Crotalinae). Molecular Phylogenetics and Evolution 49: 356-361 - get paper here
  • Guo, Peng; Qin Liu, Guanghui Zhong, Fei Zhu, Fang Yan, Ting Tang, Rong Xiao, Min Fang, Ping Wang and Xin Fu 2015. Cryptic diversity of green pitvipers in Yunnan, South-west China (Squamata, Viperidae). Amphibia-Reptilia 36 (3): 265 - 276 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Orlov, N. L., S. A. Ryabov, B. N. Thanh & H Cuc, T. 2004. A new species of Trimeresurus (Ophidia: Viperidae: Crotalinae) from Karst region in Central Vietnam. Russ. J. Herpetol. 11 (2): 139-149 - get paper here
  • Rathee, Y. S., Purkayastha, J., Lalremsanga, H. T., Dalal, S., Biakzuala, L., Muansanga, L., & Mirza, Z. A. 2022. A new cryptic species of green pit viper of the genus Trimeresurus Lacépède, 1804 (Serpentes, Viperidae) from northeast India. PloS one, 17(5), e0268402 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Ziegler, Thomas; Annemarie Ohler; Vu Ngoc Thanh; Le Khac Quyet; Nguyen Xuan Thuan; Dinh Huy Tri & Bui Ngoc Thanh 2006. Review of the amphibian and reptile diversity of Phong Nha – Ke Bang National Park and adjacent areas, central Truong Son, Vietnam. Herpetologica Bonnensis II: 247-262 - get paper here
  • ZIEGLER, THOMAS; RALF HENDRIX, VU NGOC THANH, MARTINA VOGT, BERNHARD FORSTER & DANG NGOC KIEN 2007. The diversity of a snake community in a karst forest ecosystem in the central Truong Son, Vietnam, with an identification key. Zootaxa 1493: 1-40 - get paper here
 
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