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Trimeresurus yunnanensis SCHMIDT, 1925

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Yunnan Bamboo pitviper
Chinese: 云南竹叶青蛇 
SynonymTrimeresurus yunnanensis SCHMIDT 1925: 4
Trimeresurus stejnegeri yunnanensis POPE 1935
Trimeresurus stejnegeri yunnanensis — ZHAO & ADLER 1993
Trimeresurus stejnegeri yunnanensis — WELCH 1994: 117
Trimeresurus yunnanensis — ZHAO 1995
Trimeresurus yunnanensis — GUMPRECHT et al. 2004
Trimeresurus yunnanensis — AO et al. 2004
Viridovipera yunnanensis — MALHOTRA & THORPE 2004
Viridovipera yunnanensis — GUO et al. 2006
Viridovipera yunnanensis — DAWSON et al. 2008
Trimeresurus (Viridovipera) yunnanensis — DAVID et al. 2011
Viridovipera yunnanensis — WALLACH et al. 2014: 791
Viridovipera yunnanensis — GUO et al. 2018
Trimeresurus yunnanensis — RATHEE et al. 2022
Viridovipera yunnanensis — WANG et al. 2022
Trimeresurus (Viridovipera) yunnanensis — MIRZA et al. 2023 
DistributionS China (Yunnan and southwestern Sichuan), N Myanmar, India (Meghalaya, Nagaland), elevations up to 2845 m

Type locality: Tengyueh [=Tengchiong Co.], Yunnan, China.  
Reproductionovovivparous 
TypesHolotype: AMNH 21058 
DiagnosisDIAGNOSIS (DIAGNOSTIC CHARACTERS). Scales in 19 (rarely 21) longitudinal rows at midbody and 19(–21) on neck; 9–10 (rarely 11) upper labials, first upper labials separated from nasals by a distinct suture; a single narrow supraocular, sometimes divided by transverse suture; 11–16 scales in a line between supraoculars; above bright to dark green, below pale green to whitish, the two separated by a bicolored orange or brown (below) and white (above) (males) or white only or absent (females) ventrolateral stripe, which occupies the whole of the outermost scale row and a portion of the second row; ventrals 155–165 (–170); subcaudals (58–) 61–68, all paired; hemipenes short, spinose beyond bifurcation. Total length 750 mm, tail length 145 mm. [after LEVITON 2003] 
CommentVenomous!

Habitat: primarily arboreal (Harrington et al. 2018).

Distribution: the single record from India (Meghalaya) is from 1919 (specimen in BNHS, fide GBIF). 
EtymologyNamed after the type locality. 
References
  • Ao J. M., David P., Bordoloi S., Ohler A. 2004. Notes on a collection of snakes from Nagaland, Northeast India, with 19 new records for this state. Russian Journal of Herpetology 11 (2): 155 – 162
  • David, P., G. Vogel, O.S.G. Pauwels & N. Vidal 2002. Description of a new species of the genus Trimeresurus from Thailand, related to Trimeresurus stejnegeri Schmidt, 1925 (Serpentes, Crotalidae). Natural History Journal of Chulalongkorn University, 2 (1): 5-19 - get paper here
  • DAVID, P., N. VIDAL & S. G. PAUWELS 2001. A morphological study of Stejneger's pitviper Trimeresurus stejnegeri (Serpentes, Viperidae, Crotalinae), with the description of a new species from Thailand. Russ. J. Herpetol. 8 (3): 205-222 - get paper here
  • David, Patrick, Ashok Captain and Bharat B. Bhatt 2002. On the occurrence of Trimeresurus medoensis Djao in: Djao & Jaing, 1977 (Serpentes, Viperidae, Crotalinae) in India, with a redescription of the species and notes on its biology. Hamadryad 26(2):210-226 [2001] - get paper here
  • DAVID, PATRICK; GERNOT VOGEL & ALAIN DUBOIS 2011. On the need to follow rigorously the Rules of the Code for the subsequent designation of a nucleospecies (type species) for a nominal genus which lacked one: the case of the nominal genus Trimeresurus Lacépède, 1804 (Reptilia: Squamata: Viperidae). Zootaxa 2992: 1–51 - get paper here
  • Dawson, Karen; Anita Malhotra, Roger S. Thorpe, Peng Guo, Mrinalini and Thomas Ziegler 2008. Mitochondrial DNA analysis reveals a new member of the Asian pitviper genus Viridovipera (Serpentes: Viperidae: Crotalinae). Molecular Phylogenetics and Evolution 49: 356-361 - get paper here
  • Gumprecht, A.; Tillack, F.; Orlov, N.L.; Captain, A. & Ryabow, S. 2004. Asian pitvipers. Geitje Books, Berlin, 368 pp.
  • Guo, P.; Jadin, R.C.; Malhotra, A. & Li, C. 2009. An investigation of the cranial evolution of Asian pitvipers (Serpentes: Crotalinae), with comments on the phylogenetic position of Peltopelor macrolepis. Acta Zoologica 91: 402-407 - get paper here
  • Guo, Peng; Lu, Shunqing; Huang, Song; Zhao, Hui; Zhao, Ermi 2006. Hemipenial morphology of five Asian pitvipers, with a discussion on their taxonomy. Amphibia-Reptilia 27 (1): 19-23 - get paper here
  • Guo, Peng; Malhotra, Anita; Thorpe, Roger S.; Creer, Simon; Pook, Catharine E. 2009. Comments on the systematic status of specimens belonging to the genus Viridovipera (Serpentes: Viperidae: Crotalinae) from Sichuan and Yunnan provinces of southwestern China, with a redescription of V. yunnanensis. The Herpetological Journal 19: 151-162 - get paper here
  • Guo, Peng; Qin Liu, Guanghui Zhong, Fei Zhu, Fang Yan, Ting Tang, Rong Xiao, Min Fang, Ping Wang and Xin Fu 2015. Cryptic diversity of green pitvipers in Yunnan, South-west China (Squamata, Viperidae). Amphibia-Reptilia 36 (3): 265 - 276 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Kästle , W., Rai, K. & Schleich, H.H. 2013. FIELD GUIDE to Amphibians and Reptiles of Nepal. ARCO-Nepal e.V., 625 pp. - get paper here
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  • Malhotra, Anita & Thorpe, Roger S. 2004. A phylogeny of four mitochondrial gene regions suggests a revised taxonomy for Asian pitvipers (Trimeresurus and Ovophis). Molecular Phylogenetics and Evolution 32: 83 –100 [erratum p. 680] - get paper here
  • Pope, CLIFFORD H. 1935. The Reptiles of China. Turtes, Crocodilians, Snakes, Lizards. Amer. Mus. Nat. Hist., New York, Nat. Hist. Central Asia, 10: lii, 1-604 - get paper here
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  • Rathee, Y. S., Purkayastha, J., Lalremsanga, H. T., Dalal, S., Biakzuala, L., Muansanga, L., & Mirza, Z. A. 2022. A new cryptic species of green pit viper of the genus Trimeresurus Lacépède, 1804 (Serpentes, Viperidae) from northeast India. PloS one, 17(5), e0268402 - get paper here
  • Schmidt, KARL P. 1925. New reptiles and a new salamander from China. American Museum Novitates (157): 1-5. - get paper here
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
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  • Yang, J.-H., X.-Y. Huang, J.-F. Ye, S.-P. Yang, X.-.C. Zhang & B.P.-L. Chan 2019. A report on the herpetofauna of Tengchong Section of Gaoligongshan National Nature Reserve, China. Journal of Threatened Taxa 11(11): 14434–14451 - get paper here
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