Tytthoscincus atrigularis (STEJNEGER, 1908)

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Higher Taxa	Scincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies
Common Names	E: Zamboanga Sphenomorphus
Synonym	Sphenomorphus atrigularis STEJNEGER 1908: 199 Sphenomorphus atrigularis — TAYLOR 1922 Lygosoma (Sphenomorphus) atrigularis — SMITH 1937: 221 Sphenomorphus atrigularis — MITTLEMAN 1952: 22 Sphenomorphus atrigularis — BROWN & ALCALA 1980: 158 Tytthoscincus atrigularis — LINKEM et al. 2011 Tytthoscincus atrigularis — GRISMER et al. 2016
Distribution	Philippines (Mindanao, Zamboanga Peninsula, Basilan) Type locality: Mindanao Island.
Reproduction	oviparous (phylogenetic imputation, fide Zimin et al. 2022)
Types	Holotype: USNM 37386, Datu Ahib's place, base of Malingdang Mtns, Catagan, Misamis Province, Mondanao, Philippines, collected E.A. Mearns, 24.v.1906.
Diagnosis	Diagnosis: No supranasals; fronto-nasal broadly in contact with rostral, fronto-parietals distinct; four large supraoculars; no auricular lobules; distance between tip of snout and fore leg about once and a half in distance between axil and groin; fourth toe scarcely longer than third; head and body nearly equalling length of tail; temporal region covered by scales scarcely larger than dorsals [from STEJNEGER 1908].
Comment
References	Brown, R.M. et al. 2010. Species boundaries in Philippine montane forest skinks (Genus Sphenomorphus): three new species from the mountains of Luzon and clarification of the status of the poorly known S. beyeri, S. knollmanae, and S. laterimaculatus. Scient. Pap. Nat. Hist. Mus. Univ. Kansas (42): 1-27 Brown, Rafe M., Jimmy A. McGuire, John W. Ferner and Angel C. Alcala 1999. New species of diminutive scincid lizard (Squamata: Lygosominae: Sphenomorphus) from Luzon Island, Republic of the Philippines. Copeia 1999 (2): 362-370 - get paper here Brown, WC. & A.C. ALCALA 1980. Philippine Lizards of the family Scincidae. Silliman Univ. Nat. Sci., Dumaguete City, Mon., Ser. 2: i-xi + 1-246. GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here Linkem, Charles W.; Arvin C. Diesmos, Rafe M. Brown 2011. Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163: 1217–1243 - get paper here Mittleman, M.B. 1952. A generic synopsis of the Lizards of the Subfamily Lygosominae. Smithson. misc. Collns, 117: 1-35. - get paper here Smith, M.A. 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39 (3): 213-234 Stejneger,L. 1908. Three new species of lizards from the Philippine Islands. Proc. US Natl. Mus. 34: 199-204 - get paper here Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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