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Agkistrodon piscivorus (LACÉPÈDE, 1789)

IUCN Red List - Agkistrodon piscivorus - Least Concern, LC
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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Namespiscivorus: Northern Cottonmouth
leucostoma: Western Cottonmouth
G: Wasser-Mokassinotter
G: Östliche Wasser-Mokassinotter [piscivorus]
G: Westliche Wassermokassinotter [leucostoma] 
SynonymCrotalus piscivorus LACÉPÈDE 1789: 130
Acontias leucostoma TROOST 1836: 176
Toxicophis piscivorus — BAIRD & GIRARD 1853
Toxicophis pugnax — BAIRD & GIRARD 1853
Trigonocephalus piscivorus — DUMÉRIL & BIBRON 1854: 1491
Ancistrodon piscivorus — COPE 1860: 336
Ancistrodon piscivorus var. pugnax — GARMAN 1884: plate VIII
Agkistrodon piscivorus piscivorus — GLOYD & CONANT 1943: 166
Agkistrodon piscivorus leucostoma — GLOYD & CONANT 1943: 164
Toxicophis pugnax — SCHMIDT 1954: 306
Agkistrodon piscivorous — MEHRTENS 1987: 355 (in error)
Agkistrodon piscivorus — CONANT & COLLINS 1991: 228
Agkistrodon piscivorus leucostoma — CONANT & COLLINS 1991: 229
Agkistrodon piscivorus — MCDIARMID, CAMPBELL & TOURÉ 1999: 238
Agkistrodon piscivorus piscivorus — CROTHER 2000: 56
Agkistrodon piscivorus leucostoma — CROTHER 2000: 56
Agkistrodon piscivorus leucostoma — DIXON 2000
Agkistrodon piscivorus leucostoma — TENNANT & BARTLETT 2000: 494
Agkistrodon piscivorus piscivorus — TENNANT & BARTLETT 2000: 490
Agkistrodon piscivorus piscivorus — CROTHER et al. 2012
Agkistrodon piscivorus — CROTHER et al. 2012
Agkistrodon piscivorus leucostoma — CROTHER et al. 2012
Agkistrodon piscivorus — WALLACH et al. 2014: 18
Agkistrodon piscivorus — BURBRINK & GUIHER 2014
Agkistrodon piscivorus — CROTHER et al. 2017
Agkistrodon piscivorus piscivorus — GUYER et al. 2018 
DistributionUSA (Texas, S/E Oklahoma, S Missouri, Arkansas, Louisiana, Mississippi, Kansas, Alabama, Georgia, Florida, South Carolina, E North Carolina, W Tennessee, W Kentucky, S Illinois, Indiana, SE Virginia)

piscivorus: Atlantic coastal plain south from SE Virginia west to Alabama, Georgia, Florida, North Carolina, South Carolina. Type locality: “Carolina”; Schmidt (1953 :225) proposed to restrict it to the vicinity of Charleston, South Carolina.

leucostoma: from the Mexican border (Rio Grande valley) through Texas, E Oklahoma to C Missouri and S Illinois, W Kentucky, Tennessee to Mobil Bay, Alabama. Type locality: "western district of Tennessee." Proposed emendation to "10 miles northeast of Bolivar, Hardeman County, Tennessee" by Gloyd & Conant (1943: 164), fide Schmidt (1953:225).  
Reproductionovovivparous; facultative parthenogenesis (BOOTH et al. 2012). 
TypesType: unlocated fide McDiarmid et al. 1999; unknown fide Burbrink & Guiher 2014.
Holotype: USNM 61063 (formerly 4262) [pugnax]
Neotype: CHAS 5604, adult male, designated by Gloyd and Conant, 1943 [leucostoma] 
DiagnosisDiagnosis (piscivorus): Combining characteristics for the subspecies Agkistrodon piscivorus piscivorus and Agkistrodon piscivorus leucostoma, we provide a diagnosis for A. piscivorus. The Northern Cottonmouth (A. piscivorus) is a mediumto large-bodied semi-aquatic pit viper with an average adult size of 76–114 cm and a maximum size of 188 cm (Gloyd & Conant, 1990; Conant & Collins, 1991), with a ratio of tail to total length of 0.13–0.19 in males and 0.12–0.18 in females. They possess a single anal plate, keeled dorsal scales and typically 25 midbody scale rows (range 23–27; Gloyd & Conant, 1990). Subcaudals range from 38 to 53 in males and from 42 to 53 in females, whereas ventral scales number 128–142 with no variation between sexes (Gloyd & Conant, 1990). Supralabials and infralabials range from 6 to 10 (mode 8) and 8 to 12 (mode 11), respectively and total postoculars + suboculars range from 1 to 4 (mode 3; Gloyd & Conant, 1990). A combination of geography and colour pattern distinguishes the Northern Cottonmouth from the Florida Cottonmouth. There are 10–17 dark cross-bands on an olive, brown or black background, whereas A. conanti features 11–16 crossbands. Crossbands in A. piscivorus are often indistinguishable from the ground colour in adults, yet may be prominent or subdued in adult A. conanti. The head is typically black or brown, lacking vertical rostral stripes; a dark cheek stripe is present in juveniles but may be subdued or indistinguishable from the ground colour in adults yet often present in adult A. conanti (Gloyd & Conant, 1990; Conant & Collins, 1991, BURBRINK & GUIHER 2014).


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CommentVenomous!

Taxonomy: The monophyletic Florida subspecies of A. piscivorus may be a distinct species (at 4.8% SD), whereas two western subspecies of A. contortrix also appear to constitute a single distinct species, pending additional analyses. Both species of Agkistrodon can be used as suitable ectothermic models to gauge impacts of future climate change (DOUGLAS et al. 2009). Strickland et al. 2014 found two poorly resolved clades roughly corresponding to the Florida and continental populations but their finding does not support the splitting suggested by Burbrink & Guiher 2015.

Synonymy: Burbrink & Guiher (2015) synonymized A. p. piscivorus and A. p. leucostoma into a single species, A. piscivorus. They split off A. conanti as separate species whose range completely falls into that of A. piscivorus.

ICZN opinion 1463 retains the name of LACÉPÈDE 1789.

Hybridization: Hybridization with the Florida Cottonmouth (A. conanti) occurs in the mid-Atlantic coastal plains in southern North Carolina to the southern coastal plains in south-eastern Louisiana. 
EtymologySpecific epithet refers to dietary habit, derived from Latin pisces and vorare, translated as ‘fish’ and ‘to devour’, respectively. 
References
  • Allen, Morrow J. 1932. A survey of the Amphibians and reptiles of Harrison County, Mississippi. American Museum Novitates (542): 1-20 - get paper here
  • Bailey, F.C., V.A. Cobb, T.R. Rainwater, T. Worrall & M. Klukowski 2009. Adrenocortical Effects of Human Encounters on Free-Ranging Cottonmouths (Agkistrodon piscivorus) Journal of Herpetology 43 (2): 260-266. - get paper here
  • Baird, S. F. and C. Girard. 1853. Catalogue of North American Reptiles in the Museum of the Smithsonian Institution. Part 1.-Serpents. Smithsonian Inst., Washington, xvi + 172 pp. - get paper here
  • Blem, C.R. 1997. Lipid Reserves of the Eastern Cottonmouth (Agkistrodon piscivorus) at the Northern Edge of its Range Copeia 1997 (1): 53-59. - get paper here
  • Blem, Charles R.;Blem, Leann B. 1995. The eastern cottonmouth (Agkistrodon piscivorus) at the northern edge of its range. Journal of Herpetology 29 (3): 391-398 - get paper here
  • Booth, Warren; Charles F. Smith, Pamela H. Eskridge, Shannon K. Hoss, Joseph R. Mendelson III, and Gordon W. Schuett 2012. Facultative parthenogenesis discovered in wild vertebrates. Biology Letters 8: 983-985 - get paper here
  • Booth, Warren; Gordon W. Schuett 2015. The emerging phylogenetic pattern of parthenogenesis in snakes. Biological Journal of the Linnean Society 118 (2): 172–186, DOI: 10.1111/bij.12744
  • Braswell, A.L.; Palmer, W.M. & Beane, J.C. 2003. Venomous Snakes of North Carolina. North Carolina State Museum of Natural Sciences, 31 pp.
  • Burbrink, Frank T. and Timothy J. Guiher. 2014. Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus Agkistrodon. Zoological Journal of the Linnean Society, 173: 505–526 - get paper here
  • Burkett, Ray D. 1969. Natural history of cottonmouth moccasin, Agkistrodon piscivorus (Reptilia). Univ. Kansas Publ. Mus. Nat. Hist. 17(9): 435-491
  • Burt, Charles E. 1935. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16 (3): 311-336 - get paper here
  • CAMPBELL, EMMA FELKER & JOHN B. HEWLETT. 2021. AGKISTRODON PISCIVORUS (Cottonmouth). DIET. Herpetological Review 52 (2): 416–417. - get paper here
  • Camper, Jeffrey D. 2019. The Reptiles of South Carolina. University of South Carolina Press, 288 pp. [review in Copeia 107 (3): 590, 2019] - get paper here
  • Carpenter, Charles C. 1977. Communication and Displays of Snakes. American Zoologist 17: 217-223 - get paper here
  • Chiszar, D., C. Radcliffe, R. Boyd, A. Radcliffe, H. Yun, H.M. Smith, T. Boyer, B. Atkins & F. Feiler 1986. Trailing Behavior in Cottonmouths (Agkistrodon piscivorus) Journal of Herpetology 20 (2): 269-272. - get paper here
  • Colbert, Joseph E., Kimberly M. Andrews and Terry M. Norton. 2014. Agkistrodon piscivorus (cottonmouth) diet and prey size. Herpetological Review 45 (4): 703-704 - get paper here
  • Collins, J.T. & Collins, S.L. 2009. A pocket guide to Kansas snakes, 2nd ed. Great Plains Nature Center, Wichita, 69 pp.
  • Collins, J.T. & Collins, S.L. 2010. A pocket guide to Kansas snakes, 3rd ed. Great Plains Nature Center, Wichita, 69 pp.
  • Conant, R. 1933. Three Generations of Cottonmouths, Agkistrodon piscivorus (Lacépède). Copeia, 1933(1), 43-43 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Connior, Matthew B. 2014. Geographical Distribution: Agkistrodon piscivorus leucostoma (western cottonmouth). Herpetological Review 45 (4): 662 - get paper here
  • Cope, E.D. 1900. The crocodilians, lizards and snakes of North America. Ann. Rep. U.S. Natl. Mus. 1898: 153-1270 - get paper here
  • Crabtree, Todd, Dwayne Estes and Floyd Scott 2010. Geographic distribution: Agkistrodon piscivorus leucostoma. Herpetological Review 41 (3): 378-379 - get paper here
  • Crane, Adam L. and Brian D. Greene 2008. The Effect of Reproductive Condition on Thermoregulation in Female Agkistrodon piscivorus Near the Northwestern Range Limit. Herpetologica 64 (2): 156-167 - get paper here
  • Cross, C.L. & C.E. Petersen 2001. Modeling Snake Microhabitat from Radiotelemetry Studies Using Polytomous Logistic Regression Journal of Herpetology 35 (4): 590-597. - get paper here
  • Crother, B. I. 2000. Scientific and standard English names of amphibians and reptiles of North America north of Mexico, with comments regarding confidence in our understanding. Herpetological Circular 29: 1-82
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Delisle, Z J; Delgado-Acevedo, J & Lutterschmidt, W I 2018. Agkistrodon piscivorus (Northern Cottonmouth) Predation. Herpetological Review 49 (2): 333 - get paper here
  • Delisle, Z J; Delgado-Acevedo, J & Lutterschmidt, W I 2018. Agkistrodon piscivorus (Western Cottonmouth) Predation. Herpetological Review 49 (3): 541 - get paper here
  • Delisle, Z J; Delgado-Acevedo, J; Ransom, D & Lyman-Henley, L; 2019. Agkistrodon piscivorus (Northern Cottonmouth) Reproduction/copulation following parturition. Herpetological Review 50 (1): 148 - get paper here
  • Delisle, Zackary J.; Dean Ransom, William I. Lutterschmidt & Johanna Delgado-Acevedo 2019. Multiscale Spatiotemporal Habitat Selection of Northern Cottonmouths. Journal of Herpetology 53 (3): 187-195 - get paper here
  • Dixon, James R. 2000. Amphibians and reptiles of Texas, second edition. Texas A&M University Press, 421 pp.
  • Dodge, Derec and Brian T. MIller. 2013. Geographic Distribution: Agkistrodon piscivorus leucostoma (western cottonmouth). Herpetological Review 44 (2): 274 - get paper here
  • Douglas, Michael E.;Marlis R. Douglas, Gordon W. Schuett and Louis W. Porras 2009. Climate change and evolution of the New World pitviper genus Agkistrodon (Viperidae). Journal of Biogeography 36: 1164-1180 - get paper here
  • Duméril, A. M. C., Bibron, G. & DUMÉRIL, A. H. A., 1854. Erpétologie générale ou histoire naturelle complète des reptiles. Tome septième. Deuxième partie, comprenant l'histoire des serpents venimeux. Paris, Librairie Encyclopédique de Roret: i-xii + 781-1536 - get paper here
  • Dunson, W.A. & J. Freda 1985. Water Permeability of the Skin of the Amphibious Snake, Agkistrodon piscivorus Journal of Herpetology 19 (1): 93-98. - get paper here
  • Enge, Kevin M. 2013. Geographic Distribution: Agkistrodon piscivorus conanti (Florida cottonmouth). Herpetological Review 44 (3): 475 - get paper here
  • Enge, Kevin M., Jonathan D. Mays and Bess B. Harris. 2016. Geographic Distribution: Agkistrodon piscivorus conanti (Florida Cottonmouth). Herpetological Review 47 (1): 82 - get paper here
  • Ernst, C.H., & Barbour, R.W. 1989. Snakes of eastern North America. George Mason Univ. Press, Fairfax, VA 282 pp.
  • Eversole, C B; 2019. Agkistrodon piscivorus (Cottonmouth) Prey size. Herpetological Review 50 (3): 583 - get paper here
  • Franklin, Carl J. 2006. Agkistrodon piscivorus leucostoma Diet. Herpetological Review 37 (1): 93 - get paper here
  • Friers, Josh and James P. Flaherty. 2015. Agkistrodon piscivorus conanti (Florida cottonmouth) diet. Herpetological Review 46 (4): 636-637 - get paper here
  • Garman, Samuel 1884. The reptiles and batrachians of North America. Mem. Mus. comp. Zool, Cambridge (Massachusetts), 8 (3): xxxiv + 185 pp. [1883] [CNAH reprint 10] - get paper here
  • Gibbons, J.W. & M.E. Dorcas 2002. Defensive Behavior of Cottonmouths (Agkistrodon piscivorus) toward Humans Copeia 2002 (1): 195-198. - get paper here
  • GIBBONS, PARKER W. 2021. AGKISTRODON PISCIVORUS (Cottonmouth). DEFENSIVE BEHAVIOR. Herpetological Review 52 (1): 145–146. - get paper here
  • Glaudas, Xavier; Winne, Christopher T. 2007. Do warning displays predict striking behavior in a viperid snake, the cottonmouth (Agkistrodon piscivorus)? Canadian Journal of Zoology 85.4: 574-578 - get paper here
  • Gloyd, H. and Conant, R. 1943. A synopsis of the American forms of Agkistrodon (Copperheads and Moccasins). Bull. Chicago Acad. Sci. 7: 147-170
  • Gloyd, H. K. 1969. Two additional subspecies of North American crotalid snakes, genus Agkistrodon. Proc. Biol. Soc. Washington 82: 219-232 - get paper here
  • Goodman, J.D. 1958. Material ingested by the cottonmouth, Agkistrodon piscivorus, at Reelfot Lake, Tennessee Copeia 1958 (2): 149. - get paper here
  • Graham, Sean P. 2013. How Frequently Do Cottonmouths (Agkistrodon piscivorus) Bask in Trees? Journal of Herpetology Sep 2013, Vol. 47, No. 3: 428-431. - get paper here
  • Graham, Sean P. 2015. Geographic Distribution: Agkistrodon piscivorus (cottonmouth). Herpetological Review 46 (4): 571 - get paper here
  • Graham, Sean P. and Geoffrey G. Sorrel. 2010. Agkistrodon piscivorus: Intersexual Combat. Herpetological Review 41 (3): 359-360 - get paper here
  • Grajal-Puche, Alejandro, Jillian Josimovich, Bryan Falk and Robert N. Reed. 2016. Agkistrodon piscivorus conanti (Florida Cottonmouth) Diet. Herpetological Review 47 (2): 307 - get paper here
  • Guiher, Timothy J. and Frank T. Burbrink 2008. Demographic and phylogeographic histories of two venomous North American snakes of the genus Agkistrodon. Molecular Phylogenetics and Evolution 48 (2): 543-553 - get paper here
  • Guyer, Craig; Mark A. Bailey, and Robert H. Mount 2018. Lizards and snakes of Alabama. University of Alabama Press, 397 pp. - get paper here
  • Hanscom, R. J., Higham, T. E., Ryan, D., & Clark, R. W. 2023. Ambush hunting in snakes: behaviour, function, and diversity. In: D. Penning (ed.), Snakes: morphology, function, and ecology. Hauppauge, New York: Nova Science Publishers - get paper here
  • Hecht-Karkasz, K. A., Drayer, A. N. & Hmes, J. G. 2012. Geographic distribution: Agkistrodon piscivorus (cottonmouth). Herpetological Review 43: 446-447 - get paper here
  • Herr, Mark W. and Sean P. Graham. 2014. Agkistrodon piscivorus (cottonmouth) diet. Herpetological Review 45 (3): 511 - get paper here
  • Hill, Jacques G., III and Steven J. Beaupre. 2008. Body size, growth and reproduction in a population of Western Cottonmouths Agkistrodon piscivorus leucostoma, in the Ozark Mountains of northwest Arkansas. Copeia 2008 (1): 105-114 - get paper here
  • Hill, Robert L. 2011. Geographic Distribution: Agkistrodon piscivorus piscivorus(eastern cottonmouth). Herpetological Review 42 (3): 392 - get paper here
  • Hnizdo, J. & N. Hagen 1996. Bemerkungen zur Herpetofauna der Südstaaten der USA. Sauria 18 (4): 3-10 - get paper here
  • Hoss, Shannon K.; Douglas H. Deutschman, Warren Booth and Rulon W. Clark 2015. Post-birth separation affects the affiliative behaviour of kin in a pitviper with maternal attendance. Biological Journal of the Linnean Society - get paper here
  • ICZN 1987. Opinion 1463. De Lacepede 1788-1789, Histoire Naturelle des Serpens and later editions: rejected as a non-binominal work. Bull. Zool. Nomenclature 44(4): 265-267. - get paper here
  • Irwin, Kelly J. 2004. Arkansas Snake Guide. Arkansas Game & Fish Commission, Little Rock, 50 pp.
  • Jensen, John B.; Carlos D. Camp, Whit Gibbons, & Matt J. Elliott 2008. Amphibians and reptiles of Georgia. University of Georgia Press, 575 pp.
  • Jiang Z, Castoe T, Austin C, Burbrink F, Herron M, McGuire J, Parkinson C, Pollock D 2007. Comparative mitochondrial genomics of snakes: extraordinary substitution rate dynamics and functionality of the duplicate control region. BMC Evolutionary Biology 7:123 - get paper here
  • Johnson, L.F., J.S. Jacob & P. Torrance 1982. Annual Testicular and Androgenic Cycles of the Cottonmouth (Agkistrodon piscivorus) in Alabama Herpetologica 38 (1): 16-25. - get paper here
  • Kardong, K.V. 1975. Prey Capture in the Cottonmouth Snake (Agkistrodon piscivorus) Journal of Herpetology 9 (2): 169-175. - get paper here
  • Kardong, K.V. 1977. Kinesis of the Jaw Apparatus during Swallowing in the Cottonmouth Snake, Agkistrodon piscivorus Copeia 1977 (2): 338-348. - get paper here
  • Kardong, K.V. 1982. Comparative Study of Changes in Prey Capture Behavior of the Cottonmouth (Agkistrodon piscivorus) and Egyptian Cobra (Naja haje) Copeia 1982 (2): 337-343. - get paper here
  • Keith, J., R. Lee & D. Chiszar 1985. Spatial Orientation by Cottonmouths (Agkistrodon piscivorus) after Detecting Prey Bull. Maryland Herpetol. Soc., 21(4): 145-149. - get paper here
  • Kuhns, A R; Phillips, C A; Dreslik, M J; Corcoran, M & Crawford, J A 2018. Agkistrodon piscivorus (Cottonmouth) Diet. Herpetological Review 49 (2): 332-333 - get paper here
  • Kwet, Axel 2015. Neue Mokassinschlangen in Nordamerika. Terraria-Elaphe 2015 (5): 54-55 - get paper here
  • Lacepède, B. G. E. 1789. Histoire Naturelle des Quadrupèdes Ovipares et de Serpens. Vol.2. lmprimerie du Roi, Hôtel de Thou, Paris, 671 pp. - get paper here
  • Laita, Mark 2013. Serpentine. Abrams and PQ Blackwell, Auckland, New Zealand, 200 unnumbered pages
  • Lee, James R., Lynn McCoy and Annie Walker. 2014. Agkistrodon piscivorus (cottonmouth) diet. Herpetological Review 45 (4): 703 - get paper here
  • Lee, John R. 1996. Agkistrodon piscivorus piscivorus (Eastern cottonmouth). Coloration. Herpetological Review 27 (1): 22 - get paper here
  • Lillywhite, H. B.; J. B. Pfaller and C. M. Sheehy III 2015. Feeding preferences and responses to prey in insular neonatal Florida cottonmouth snakes. Journal of Zoology, DOI: 10.1111/jzo.12256 - get paper here
  • Lillywhite, Harvey B. and Coleman M. Sheehy III 2019. The Unique Insular Population of Cottonmouth Snakes at Seahorse Key. in: Lillywhite, H.B & Martins, M. (eds.) Islands and Snakes, OUP, pp. 201 ff.
  • Linzey, D.W., & Clifford, M.J. 1981. Snakes of Virgina. Univ. Press of Virginia, Charlottesville 159 pp.
  • Lönnberg, Einar 1894. Notes on reptiles and batrachians collected in Florida in 1892 and 1893. Proc. US Natl. Mus. 17 (1003): 317-339 - get paper here
  • Love, B. 2011. WESTERN HERP PERSPECTIVES: Nicht ausreichend gewürdigte Reptilien. Reptilia (Münster) 16 (88): 14 - get paper here
  • LUTTERSCHMIDT, WILLIAM I.; ZANDER E. PERELMAN, DREW R. NEYLAND, MONTE L. THIES & ERIC D. ROTH. 2021. AGKISTRODON PISCIVORUS (Northern Cottonmouth). DIET. Herpetological Review 52 (3): 655–656. - get paper here
  • Martin, D.L. 1984. An Instance of Sexual Defense in the Cottonmouth, Agkistrodon piscivorus Copeia 1984 (3): 772-774. - get paper here
  • McAllister, C. T. 2017. Agkistrodon piscivorus leucostoma (Western Cottonmouth) Diet. Herpetological Review 48 (2): 443-444 - get paper here
  • McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp.
  • Merkle, Donald A. 1985. Genetic variation in the Eastern cottonmouth, Agkistrodon piscivorus piscivorus (Lacépède) (Reptilia: Crotalidae) at the northern edge of its range. Brimleyana 11: 55-61 - get paper here
  • Mitchell, J. C. & Reay, K.K. 1999. Atlas of amphibians and reptiles in Virginia. Specialty Publication 1, VA Dept of Game and Fisheries, 122 pp. - get paper here
  • Mitchell, J.C. 1994. The reptiles of Virginia. Virginia Department of Game and Inland Fisheries, ca. 350 pp.
  • Neal, Trey and David A. Steen. 2015. Agkistrodon piscivorus (cottonmouth) predation. Herpetological Review 46 (2): 264 - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Palis, J. G. 2016. Snakes of “Snake Road”. Bull. Chicago Herp. Soc. 51: 1 - get paper here
  • Palis, J. G. 2017. Agkistrodon piscivorus (Cottonmouth) Diet. Herpetological Review 48 (2): 443 - get paper here
  • Palis, John G. 2014. Agkistrodon piscivorus (cottonmouth) fire mortality. Herpetological Review 45 (4): 704-705 - get paper here
  • PALIS, JOHN G. 2022. AGKISTRODON PISCIVORUS (Northern Cottonmouth). PREDATION. Herpetological Review 53 (3): 503–504.
  • Palmer, W.M. & Braswell, A.L. 1995. Reptiles of North Carolina. Univ. North Carolina Press
  • Parkinson,C.L. 1999. Molecular systematics and biogeographical history of pitvipers as determined by mitochondrial ribosomal DNA sequences. Copeia 1999 (3): 576-586 - get paper here
  • PAUL, LAURENCE & ROBERT W. MENDYK. 2021. Glow and Behold: Biofluorescence and New Insights on the Tails of Pitvipers (Viperidae: Crotalinae) and Other Snakes. Herpetological Review 52 (2): 221–237. - get paper here
  • Perry, David A. 2013. Virginia Herpetological Society Survey of Back Bay NWR and False Cape State Park. Catesbeiana 33 (2): 48-61 - get paper here
  • Perry, J. 1978. An Observation of ”Dance” Behavior in the Western Cottonmouth, Agkistrodon piscivorus leucostoma (Reptilia, Serpentes, Viperidae) Journal of Herpetology 12 (3): 429-431. - get paper here
  • Pitts, Steven L. 2015. Agkistrodon piscivorus (cottonmouth) cold tolerance. Herpetological Review 46 (3): 441 - get paper here
  • Preston, Robert, Jr. and Dirk J. Stevenson. 2011. Geographic Distribution: Agkistrodon piscivorus (cottonmouth). Herpetological Review 42 (3): 392 - get paper here
  • Ramsey, L.W. 1948. Combat Dance and Range Extension of Agkistrodon piscivorus leucostoma Herpetologica 4 (6): 228. - get paper here
  • Rieser, Jennifer M.; Tai-De Li, Jessica L. Tingle, Daniel I. Goldman, Joseph R. Mendelson 2021. Functional consequences of convergently evolved microscopic skin features on snake locomotion. Proceedings of the National Academy of Sciences Feb 2021, 118 (6) e2018264118; DOI: 10.1073/pnas.2018264118 - get paper here
  • Roth, E.D. 2005. Spatial Ecology of a Cottonmouth (Agkistrodon piscivorus) Population in East Texas Journal of Herpetology 39 (2): 308-312 - get paper here
  • Rush, Scott A.; Kim Sash, John Carroll, Bill Palmer, and Aaron T. Fisk 2014. Feeding Ecology of the Snake Community of the Red Hills Region Relative to Management for Northern Bobwhite: Assessing the Diet of Snakes Using Stable Isotopes. Copeia Jun 2014, Vol. 2014, No. 2: 288-296. - get paper here
  • Schmidt, D. & Kunz, K. 2005. Ernährung von Schlangen. Natur und Tier Verlag, Münster, 159 pp. - get paper here
  • Schmidt, Karl P. 1954. Omissions from the Sixth Edition of the Check List of North American Amphibians and Reptiles. Copeia 1954: 304-306 - get paper here
  • Scott, D.E., R.U. Fischer, J.D. Congdon & S.A. Busa 1995. Whole Body Lipid Dynamics and Reproduction in the Eastern Cottonmouth, Agkistrodon piscivorus Herpetologica 51 (4): 472-487. - get paper here
  • Sheehy, C. M., III, et al. 2017. Cannibalism and changing food resources in insular cottonmouth snakes. Herpetological Review 48 (2): 310-312 - get paper here
  • Siegel, Dustin S.; David M. Sever, Justin L. Rheubert, and Kevin M. Gribbins 2010. REPRODUCTIVE BIOLOGY OF AGKISTRODON PISCIVORUS LACÉPÈDE (SQUAMATA, SERPENTES, VIPERIDAE, CROTALINAE). Herpetological Monographs 23 (1): 74-107[2009] - get paper here
  • Snyder, Richard C. 1945. Notes on the snakes of southeastern Alabama. Copeia 1945 (3): 173-174 - get paper here
  • Spinner, L. 2017. Auf der Lauer: Wassermokassinottern in Natur und Terrarium. Reptilia (Münster) 22 (124): 34-40 - get paper here
  • Steehouder, A.M. 1988. Agkistrodon piscivorus, the Cottonmouth. Litteratura Serpentium 8 (4): 173-181 - get paper here
  • Strickland, Jason L.; Christopher L. Parkinson, J. Kelly McCoy, and Loren K. Ammerman 2014. Phylogeography of Agkistrodon piscivorus with Emphasis on the Western Limit of Its Range. Copeia Dec 2014, Vol. 2014, No. 4: 639-649. - get paper here
  • Strimple, Pete 1994. Maintenance, feeding, and growth of a juvenile Eastern cottonmouth, Agkistrodon piscivorus piscivorus, during its first eighteen months in captivity. Litteratura Serpentium 14 (3): 70-73 - get paper here
  • Strimple, Pete 1998. The cottonmouth or water moccasin Agkistrodon piscivorus (Lacepède), part 3. Litteratura Serpentium 18 (6): 146-153 - get paper here
  • Strimple, Pete 1998. The cottonmouth or water moccasin Agkistrodon piscivorus (Lacepède), part 2. Litteratura Serpentium 18 (5): 126-132 - get paper here
  • Strimple, Pete 1998. The cottonmouth or water moccasin Agkistrodon piscivorus (Lacepède), part 1. Litteratura Serpentium 18 (4): 99-103 - get paper here
  • Strimple, Pete 1999. The cottonmouth or watermocassin, Agkistrodon piscivorus, deel 4. Litteratura Serpentium 19 (1): 16-27 - get paper here
  • Tennant, A. 2003. Snakes of North America - Eastern and Central Regions, revised edition. Lone Star Books, 605 pp.
  • Tennant, A. & Bartlett, R.D. 2000. Snakes of North America - Eastern and Central Regions. Gulf Publishing, Houston, TX, 588 pp.
  • Tetzlaff, Sasha J., David L. Tetzlaff, Michelle Schoenbeck and Nick Clark. 2014. Agkistrodon piscivorus conanti (Florida cottonmouth) diet and phenotype. Herpetological Review 45 (3): 511-512 - get paper here
  • Troost 1836. On a new genus of serpents, and two new species of the genus Heterodon, inhabiting Tennessee. Ann. Lyc. Nat. Hist. New York 3, p. 174-190 - get paper here
  • Trutnau, L. 1984. Ein Beitrag zur Kenntnis des Kupferkopfs - Agkistrodon contortrix (LINNAEUS 1766). Herpetofauna 7 (35): 14-26 - get paper here
  • Trutnau, L. 1997. Zur Biologie, Haltung und Nachzucht der Wassermokassinotter Agkistrodon piscivorus (Lacépède, 1789). Herpetofauna 19 (108): 25-34 - get paper here
  • Turner, Jeff 2019. Agkistrodon piscivorus piscivorus (Northern Cottonmouth). Catesbeiana 39 (1): 31 - get paper here
  • Vaughn, R. Kathryn;Baumgardner, George D. 1996. Geographic Distribution. Agkistrodon piscivorus leucostoma. Herpetological Review 27 (1): 33 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Weidler, J.M., Thies, M.L., Smith-Herron, A., Tivador, E.J., III, Roth, E.D. & Lutterschmidt, W.I. 2017. Agkistrodon piscivorus leucostoma (Western Cottonmouth) Unusual field injury. Herpetological Review 48 (3): 664-665. - get paper here
  • Werler, John E. & James R. Dixon 2000. Texas Snakes. University of Texas Press, 544 pages
  • Wharton, C. H. 1969. The cottonmouth moccasin on sea horse key, Florida. Bulletin of the Florida State Museum 14 (3): 227-272 - get paper here
  • Wharton, C.H. 1960. Birth and Behavior of a Brood of Cottonmouths, Agkistrodon piscivorus piscivorus with Notes on Tail-Luring Herpetologica 16 (2): 125-129. - get paper here
  • Wharton, C.H. 1966. Reproduction and Growth in the Cottonmouths, Agkistrodon piscivorus Lacépède, of Cedar Keys, Florida Copeia 1966 (2): 149-161. - get paper here
  • Zaidan, F., III, D.L. Kreider & S.J. Beaupre 2003. Testosterone cycles and reproductive energetics: Implications for northern range limits of the cottonmouth (Agkistrodon piscivorus leucostoma) Copeia 2003 (2): 231-240. - get paper here
 
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