| Diagnosis | Diagnosis: Anolis heterodermus shares short limbs, a large, casqued head, a prehensile tail without caudal autotomy, and lamellar subdigital scales of all digits extending from the most proximal phalanges with the other species of the Phenacosaurus clade. Anolis heterodermus differs from A. proboscis, A. orcesi, A. euskalerriari and A. nicefori by the presence of granular scales surrounding dorsal heterogeneous flat scales. From A. richteri, A. tequendama sp. nov., A. tetarii, A. vanzolinii, A. inderenae and A. quimbaya sp. nov., by the presence of a unicoloured purple, red, or black dewlap and a U-shaped crown. Anolis heterodermus further differs from A. richteri, A. tequendama sp. nov., A. tetarii, A. vanzolinii, and A. quimbaya sp. nov., by the presence of a continuous row of enlarged sublabials reaching the mouth commissure; from A. vanzolinii, A. tequendama sp. nov. and A. quimbaya sp. nov. by a continuous nuchal crest; and from A. vanzolinii by < 24 expanded lamellae under second and third phalange of fourth toe and < 35 lamellae in total fourth toe. (Moreno-Arias et al. 2023) |
| Comment | Type species: Anolis heterodermus DUMÉRIL 1851 is the type species of the genus Phenacosaurus BARBOUR 1920. Phenacosaurus was synonymized with Anolis by HASS et al. 1993.
Key: Moreno-Arias et al. 2023 provide a key to the 11 species of the Phenacosaurus clade of Anolis.
Synonymy: Lazell 1969 synonymized A. richteri with A. heterodermus. Moreno-Arias et al. 2023 revalidated A. richteri.
Species groups: Dactyloa heteroderma species group (fide NICHOLSON et al. 2012). Prates et al. 2019 redefined the heterodermus series as follows: The largest crown clade that includes A. heterodermus and excludes A. aequatorialis, A. latifrons, A. nasofrontalis, A. punctatus, and A. roquet. The Phenacosaurus clade of Anolis includes the following 11 species: euskalerriari, heterodermus, inderenae, nicefori, orcesi, proboscis, richteri, tetarii, tequendama, quimbaya, vanzolinii fide Moreno-Arias et al. 2023 (although other species have been assigned to Phenacosaurus as well) |
| References |
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