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Ctenotus grandis STORR, 1969

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
SubspeciesCtenotus grandis grandis STORR 1968
Ctenotus grandis titan STORR 1980 
Common NamesE: Grand Ctenotus 
SynonymCtenotus grandis STORR 1969: 100
Ctenotus grandis — COGGER 1983: 146
Minervascincus grandis — WELLS & WELLINGTON 1985: 34
Ctenotus grandis — COGGER 2000: 426
Ctenotus grandis — WILSON & SWAN 2010

Ctenotus grandis titan STORR 1980
Minervascincus titan — WELLS & WELLINGTON 1985: 35
Ctenotus grandis titan — STORR et al. 1999 
DistributionAustralia (Northern Territory, South Australia?, Western Australia)

grandis: Tanami, Great Sandy, Gibson and Great Victoria Deserts.
titan: Pilbara and adjacent regions, Western Australia.

Type locality: 24 mi ENE of Lavation, in 28° 31’ S, 122'45’ E, W. A.  
Reproductionoviparous 
TypesHolotype: WAM R30005
Holotype: WAM R61444, from 1 km S of Exmouth, W. A. [titan] 
DiagnosisDiagnosis: A very large Ctenotus with relatively stout body and short, thick appendages; dark dorsal stripes on a greenish ground (juveniles) or reddish ground (adults); sides dark with small white spots tending to be arranged in vertical bars; second loreal usually high, pentagonal, with angular apex. (Storr 1969) 
CommentAbundance: common, with more than 500 specimens collected (Pianka 2011)

Limb morphology: 5 digits, 5 toes (Singhal et al. 2018, Cogger 2014) 
EtymologyThe specific epithet, grandis, is a Latin adjective, meaning ‘big’ or ‘large’, in reference to the larger body size of this species compared to congeners. 
References
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Couper, P., Covacevich, J., Amey, A. & Baker, A. 2006. The genera of skinks (Family Scincidae) of Australia and its island territories: diversity, distribution and identification. in: Merrick, J.R., Archer, M., Hickey, G.M. & Lee, M.S.Y. (eds.). Evolution and Zoogeography of Australasian Vertebrates. Australian Scientific Publishing, Sydney, pp. 367-384
  • Ellis, Ryan J. 2015. Ctenotus grandis grandis (grand ctenotus) birfurcation. Herpetological Review 46 (2): 256 - get paper here
  • Pianka, E. R. 1969. Sympatry of desert lizards (Ctenotus) in Western Australia. Ecology 50: 1012-1030 - get paper here
  • Pianka, E. R. 1972. Zoogeography and speciation of Australian desert lizards: an ecological perspective. Copeia 1972 (1): 127-144 - get paper here
  • Punzo, Fred & Sonia Madragon 2002. Spatial learning in Australian skinks of the genus Ctenotus (Scincidae). Amphibia-Reptilia 23 (2): 233-238 - get paper here
  • Singhal, Sonal; Huateng Huang, Maggie R. Grundler, María R. Marchán-Rivadeneira, Iris Holmes, Pascal O. Title, Stephen C. Donnellan, and Daniel L. Rabosky 2018. Does Population Structure Predict the Rate of Speciation? A Comparative Test across Australia’s Most Diverse Vertebrate Radiation. The American Naturalist - get paper here
  • Storr G M 1969. The genus Ctenotus (Lacertilia: Scincidae) in the Eastern Division of Western Australia. J. Royal Soc. Western Australia 51: 97-109 - get paper here
  • Storr G M 1970. The genus Ctenotus (Lacertilia: Scincidae) in the Northern Territory. J. Royal Soc. Western Australia 52: 97-108 [1969] - get paper here
  • Storr G M 1980. The Ctenotus grandis species-group (Lacertilia: Scincidae). Rec. West. Aust. Mus. 8 (3): 415-422 - get paper here
  • Storr, G. M. 1975. The genus Ctenotus (Lacertilia: Scincidae) in the Kimberley and North-west Divisions of Western Australia. Rec. West. Aust. Mus. 3: 209-243 - get paper here
  • Storr, G. M., Smith, L. A. & Johnstone, R. E. 1981. Lizards of Western Australia. I. Skinks. Perth: University of Western Australia Press and Western Australian Museum, 200 pp.
  • Storr, G. M.; L. A. Smith, and R. E. Johnstone 1999. Lizards of Western Australia. I. Skinks. Revised Edition. Western Australian Museum
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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