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Darevskia mixta (MÉHELY, 1909)

IUCN Red List - Darevskia mixta - Near Threatened, NT
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Higher TaxaLacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesG: Bastardeidechse 
SynonymLacerta mixta MÉHELY 1909: 581
Lacerta composita MÉHELY 1909 (nomen oblitum)
Lacerta saxicola mixta LANTZ & CYREN 1936
Lacerta (Zootoca) “mixta” — BISCHOFF 1978
Lacerta mixta — ENGELMANN et al 1993
Lacerta mixta — FU & MURPHY 1997
Darevskia mixta — ARRIBAS 1997
Lacerta mixta — FU, MURPHY & DAREVSKY 2000
Darevskia mixta — MURPHY et al. 2000
Archaeolacerta (Caucasilacerta) mixta — SINDACO et al. 2000
Lacerta mixta — SZCZERBAK 2003
Darevskia mixta — BISCHOFF 2007
Lacerta (Darevskia) mixta — SINDACO & JEREMČENKO 2008 
DistributionSW Republic of Georgia (Small Caucasus)

Type locality: Abastumani, Republic of Georgia  
Reproductionoviparous 
TypesLectotype: SMF 12087, designated by R. Mertens 1967. 
Diagnosis 
CommentSynonymy partly after KHALIKOV & ANANJEVA (pers. comm.). This species is not listed in the Zoological Record (1909) where other species described by MÉHELY are listed.

Distribution: also in Turkey (fide web site Türkiye'de Kaç Tür KertenkeleYas¸ıyor), but not in Turkey fide Gabelaia et al. 2015. 
References
  • Abramjan, A., Arakelyan, M., & Frynta, D. 2020. Does reproductive mode affect sexually selected colouration? Evaluating UV-blue spots in parthenogenetic and bisexual lizards of the genus Darevskia. Current Zoology - get paper here
  • Bischoff, W. 1978. Beiträge zur Kenntnis der Echsen des Kaukasus. Salamandra 14 (4): 178-202 - get paper here
  • Bischoff, W. 2007. Auf der Suche nach Gebirgseidechsen in drei Erdteilen. Draco 7 (27): 54-73 - get paper here
  • Darevsky,I.S. 1957. Systematics and ecology of rock lizards (Lacerta saxicola Eversmann) in Armenia [in Russian]. Zool. sb. AN Armenia SSR 10: 27-57 - get paper here
  • Doronin, Igor V. 2018. The Taxonomic Status of Lacerta mixta Méhely, 1909 (Squamata: Sauria: Lacertidae). Russ. J. Herpetol. 25 (2): 143-146 - get paper here
  • Ebel, R., Müller, J., Ramm, T. et al. 2020. First evidence of convergent lifestyle signal in reptile skull roof microanatomy. BMC Biol 18, 185 (2020) - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Freitas, S., Westram, A.M., Schwander, T., Arakelyan, M., Ilgaz, Ç., Kumlutas, Y., Harris, D.J., Carretero, M.A. and Butlin, R.K. 2022. Parthenogenesis in Darevskia lizards - a rare outcome of common hybridisation, not a common outcome of rare hybridisation Parthenogenesis, gene flow and hybridisation. Evolution - get paper here
  • Fu J.; Murphy R.W.; Darevsky I.S. 1999. Limited genetic variation in Lacerta mixta and its parthenogenetic daughter species: evidence from cytochrome b and ATPase 6 gene DNA sequences. Genetica 105 (3): 227-231
  • Fu, JINZHONG AND ROBERT W. MURPHY 1997. Toward the phylogeny of caucasian rock lizards: implications from mitochondrial DNA gene sequences (Reptilia: Lacertidae). Zoological Journal of the Linnean Society (1997), 121: 463–477. - get paper here
  • Gabelaia, Mariam, David Tarkhnishvili and Marine Murtskhvaladze. 2015. Phylogeography and morphological variation in a narrowly distributed Caucasian rock lizard, Darevskia mixta. Amphibia-Reptilia 36 (1): 45-54 - get paper here
  • Gabelaia, Mariam; Dominique Adriaens, David Tarkhnishvili 2017. Phylogenetic signals in scale shape in Caucasian rock lizards (Darevskia species). Zoologischer Anzeiger 268: 32–40 - get paper here
  • Grechko VV, Ciobanu DG, Darevsky IS, Kosushkin SA, Kramerov DA. 2006. Molecular evolution of satellite DNA repeats and speciation of lizards of the genus Darevskia (Sauria: Lacertidae). Genome 49(10):1297-307. - get paper here
  • Lantz,L.A. & Cyrén,O. 1936. Contribution à la connaissance de Lacerta saxicola EVERSMANN. Bull. Soc. zool. France (Paris) 61: 159-181 - get paper here
  • Méhely, Lajos von 1909. Materialien zu einer Systematik und Phylogenie der muralis-ähnlichen Lacerten. Annales historico-naturales Musei nationalis Hungarici, Budapest, 7 (2): 409-621 - get paper here
  • Murphy, R.W.; Fu,J.; MacCulloch, R.D.; Darevsky,I.S. & Kupriyanova,L.A. 2000. A fine line between sex and unisexuality: the phylogenetic constraints on parthenogenesis in lacertid lizards. Zool. J. Linnean Soc. 130: 527-549 - get paper here
  • Petrosyan, V., Osipov, F., Bobrov, V., Dergunova, N., Omelchenko, A., Varshavskiy, A., ... & Arakelyan, M. 2020. Species Distribution Models and Niche Partitioning among Unisexual Darevskia dahli and Its Parental Bisexual (D. portschinskii, D. mixta) Rock Lizards in the Caucasus. Mathematics, 8(8): 1329 - get paper here
  • Petrosyan, Varos; Fedor Osipov, Vladimir Bobrov, Natalia Dergunova, Evgeny Nazarenko, Andrey Omelchenko, Felix Danielyan & Marine Arakelyan 2019. Analysis of geographical distribution of the parthenogenetic rock lizard Darevskia armeniaca and its parental species (D. mixta, D. valentini) based on ecological modelling. Salamandra 55 (3): 173-190 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Alberto Venchi, Giuseppe M. Carpaneto, Marco A. Bologna 2000. The Reptiles of Anatolia: a Checklist and Zoogeographical analysis. Biogeographia, 21-2000: 441-554. - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • Tarkhnishvili, D., Gabelaia, M. and Adriaens, D. 2020. Phenotypic divergence, convergence and evolution of Caucasian rock lizards (Darevskia). Biological Journal of the Linnean Society - get paper here
  • Tuniyev B.S., Tuniyev S.B., Avcı A., Ilgaz Ç. 2014. Herpetological studies in Eastern and North-Eastern Turkey. Current Studies in Herpetology 14 (1/2): 44–53. Резюме - get paper here
  • Tuniyev, B.S. 1990. On the Independence of the Colchis Center of Amphibian and Reptile Speciation. Asiatic Herpetological Research 3: 67-84 - get paper here
  • Uzzell, T. and Darevsky, I. S. 1973. Electrophoretic examination of Lacerta mixta - a possible hybrid species (Sauria, Lacertidae). Journal of Herpetology 7 (1): 11-15. - get paper here
  • Uzzell, T. and Darevsky, I. S. 1975. Biochemicai evidence for the hybrid origin of the parthenogenetic species of the Lacerta saxicola complex (Sauria: Lacertidae), with a dicussion of some ecological and evolutionary implications. Copeia 1975 (2): 204-222. - get paper here
 
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