Hebius concelarus (MALNATE, 1963)

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Higher Taxa	Colubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names
Synonym	Amphiesma pryeri concelara MALNATE 1963 Amphiesma pryeri concelarum MALNATE 1963 Natrix pryeri — MAKI 1931 Amphiesma pryeri — MALNATE 1960: 52 (part.) Amphiesma pryeri concelarum — MALNATE 1963: 2 Amphiesma concelarum — OTA and IWANAGA 1997 Hebius concelarum — GUO et al. 2014 Amphiesma concelarum — WALLACH et al. 2014: 29 Hebius concelarus — KAITO et al. 2016
Distribution	Japan (Ryukyu Islands) Type locality: “Miyako-shima” (Miyakojima), Miyako Group, Riukiu (Ryukyu) Islands.
Reproduction	oviparous
Types	Holotype: CAS 21905; adult female; paratype CAS, malee
Diagnosis	Diagnosis: A form of Amphiesma pryeri distinguished by a reduced number of ventrals (159-165); latero-ventral black spots strongly developed; narrow, black-edged, light vertical bar antriorly on the sides of the body; head dark, the light supralabials heavily marbled with dark gray or black. (Malnate 1963) Additional details (36 characters) are available for collaborators and contributors. Please contact us for details.
Comment	Raised to species status by Ota & Iwanaga 1997. This species is classified as “vulnerable” in Japan (Ota 2000).
References	GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here Kaito, Takuma; and Mamoru Toda 2016. The biogeographical history of Asian keelback snakes of the genus Hebius (Squamata: Colubridae: Natricinae) in the Ryukyu Archipelago, Japan. Biological Journal of the Linnean Society, DOI: 10.1111/bij.12726 - get paper here Malnate, E. V. 1963. A new race of Amphiesma pryeri (Serpentes: Natricinae) from the southern Riukiu Island of Miyako-Shima. Notulae Naturae 360: 1-6 Ota, H. 2000. Current status of the threatened amphibians and reptiles of Japan. Popul. Ecol. 42: 5-9 - get paper here Ota, HIDETOSHI & SETSUKO IWANAGA 1997. A systematic review of the snakes allied to Amphiesma pryeri (Boulenger) (Squamata: Colubridae) in the Ryukyu Archipelago, Japan. Zoological Journal of the Linnean Society 121 (3): 339-360 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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