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Lygosoma boehmei ZIEGLER, SCHMITZ, HEIDRICH, VU & NGUYEN, 2007

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Higher TaxaScincidae, Lygosominae (Lygosomini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymLygosoma boehmei ZIEGLER, SCHMITZ, HEIDRICH, VU & NGUYEN 2007
Riopa boehmei — BOBROV & SEMENOV 2008
Lygosoma boehmei — GEISSLER et al. 2012
Lygosoma boehmei — LIVIGNI 2013: 279 
DistributionVietnam (Quang Binh)

Type locality: karst forest of Cha Noi, 350-400 m elevation, Phong Nha – Ke Bang National Park, Quang Binh Province, Vietnam.  
Reproduction 
TypesHolotype: ZFMK 86359: female, collected by Astrid Heidrich and Thomas Ziegler at the end of the dry season (21 June) 2006. 
DiagnosisDIAGNOSIS: The new species can be distinguished from any other Lygosoma / Riopa currently known by the following combination of characters: (1) Body elongate (SVL 86.0 mm), the distance between axilla and groin being slightly more than 3 times the length of the forelimb; (2) forelimb and hindlimb short, pentadactyl, fingers and toes widely separated when adpressed, the distance between them corresponding to the length of the hindlimb; (3) dorsum reddish brown in life, brownish grey in preservative, the surfaces of the limbs and tail being brownish black; ventral side yellowish beige to greyish below the tail in preservative, with some indistinct dark marbling; in life, chin and throat are light orange and the remaining ventral side of the body cream to light brownish, turning to greyish at the underside of the tail; sides of body and neck with irregular dark flecking; sutures of anterior supra- and infralabials edged by greyish black; (4) rostral separated from undivided frontonasal by supranasals (5) prefrontals widely separated; (6) parietals forming a suture behind interparietal, no enlarged nuchals; (7) four supraoculars; bordered by six supraciliaries; posteriorly, the fourth supraocular is bordered by a small scale (postsupraocular); (8) lower eyelid scaly; (9) two loreals; (10) seven supralabials, fifth right below the eye; (11) seven infralabials; (12) postmental undivided; (13) first pair of chinshields in broad contact, bordered by six scales; (14) ear opening small, ovoid to roundish; (15) 32 midbody scale rows; (16) 66 middorsal (paravertebral) scales (from the posterior end of parietals to insertion of hindlimb); (17) dorsal and dorsolateral scales notched, appearing as three longitudinal (pseudo-)keels on neck, body, and anterior half of tail; (18) ventral scales smooth, arranged in 81 transverse rows from first gular row between third pair of chinshields to preanals; (19) six slightly enlarged preanals; (20) 108 median subcaudal scales, not enlarged, smooth; (21) fourth toe with 14 keeled subdigital lamellae. 
CommentL. boehmei is molecularly most closely related to L. quadrupes and L. koratense.

Abundance: only known from the type locality (Meiri et al. 2017). This is one of the species called 'lost' and 'rediscovered' by Lindken et al. 2024. 
Etymologynamed after Wolfgang Böhme, German herpetologist. 
References
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • GEISSLER, PETER; TIMO HARTMANN & THY NEANG 2012. A new species of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae) from northeastern Cambodia, with an updated identification key to the genus Lygosoma in mainland Southeast Asia. Zootaxa 3190: 56–68 - get paper here
  • Geissler, Peter; Truong Quang Nguyen, Trung My Phung, Robert Wayne Van Devender, Timo Hartmann, Balázs Farkas, Thomas Ziegler and Wolfgang Böhme 2011. A review of Indochinese skinks of the genus Lygosoma Hardwicke & Gray, 1827 (Squamata: Scincidae), with natural history notes and an identification key. Biologia 66 (6): 1159-1176 - get paper here
  • Lindken T.; Anderson, C. V., Ariano-Sánchez, D., Barki, G., Biggs, C., Bowles, P., Chaitanya, R., Cronin, D. T., Jähnig, S. C., Jeschke, J. M., Kennerley, R. J., Lacher, T. E. Jr., Luedtke, J. A., Liu, C., Long, B., Mallon, D., Martin, G. M., Meiri, 2024. What factors influence the rediscovery of lost tetrapod species? Global Change Biology, 30: 1-18 - get paper here
  • LiVigni, F. (ed.) 2013. A Life for Reptiles and Amphibians, Volume 1. Chimaira, Frankfurt, 495 pp. - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Thompson, C. & Thompson, T. 2008. First contact in the Greater Mekong - new species discoveries. WWF, 40 pp. - get paper here
  • Ziegler, T. 2007. Field surveys and collection management as basis for herpetodiversity research and nature conservation in Vietnam. Proc. Int. Conf. “Development of Hochiminh City Museum of Natural History”, Ho Chi Minh City People’s committee, pp. 230-248
  • Ziegler, T; Schmitz, A; Heidrich, A; Vu, NT; Nguyen, QT 2007. A new species of Lygosoma (Squamata: Sauria: Scincidae) from the Central Truong Son, Vietnam, with notes on its molecular phylogenetic position. Revue Suisse de Zoologie 114 (2): 397-415 - get paper here
 
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