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Meroles squamulosus (PETERS, 1854)

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesE: Savanna Lizard, Common Rough-scaled (Sand) Lizard 
SynonymIchnotropis squamulosa PETERS 1854: 617
Ichnotropis squamulosa — FITZSIMONS 1943: 350
Ichnotropis squamulosa — HUEY & PIANKA 1981
Ichnotropis squamulosa — AUERBACH 1987: 130
Ichnotropis squamulosa — BROADLEY & HOWELL 1991: 18
Ichnotropis squamulosa — PIETERSEN et al. 2013
Meroles squamulosa — ENGLEDER et al. 2013
Meroles squamulosa — EDWARDS et al. 2013
Meroles squamulosus — SPAWLS et al 2018: 20
Meroles squamulosus — SPAWLS et al 2018: 203 
DistributionTanzania, Zambia, S Angola, Zimbabwe, Botswana, Namibia, N South Africa, Malawi.

Type locality: Tete, Mozambique  
Reproductionoviparous 
TypesSyntypes: ZMB 6614, 6615, 6616 
Diagnosis 
CommentSynonymy: EDWARDS et al. 2013 proposed the name Meroles squamulosa as “comb. nov.” but Engleder’s paper was published earlier. 
References
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  • Baeckens, Simon, Jan Scholliers and Raoul Van Damme. 2016. Geographic Distribution: Meroles squamulosus (Common Rough-scaled lizard). Herpetological Review 47 (1): 81 - get paper here
  • Baptista NL, António T, Branch WR. 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2) [Special Section]: 96–130 (e203) - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
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  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1962. On some reptile collections from the North-Western and North-Eastern Districts of Southern Rhodesia 1958-1961, with descriptions of four new lizards. Occ. Pap. Nat. Mus. South. Rhodesia 26 (B): 787-843
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  • Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J, Júlio T, Neef G. 2022. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles. Amphibian & Reptile Conservation 16(2): 181–214 (e322) - get paper here
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  • EDWARDS, SHELLEY; WILLIAM R. BRANCH, BIEKE VANHOOYDONCK, ANTHONY HERREL, G. JOHN MEASEY, KRYSTAL A. TOLLEY 2013. Taxonomic adjustments in the systematics of the southern African lacertid lizards (Sauria: Lacertidae). Zootaxa 3669 (2): 101–114 - get paper here
  • Engleder, A., Haring, E., Kirchhof, S. and Mayer, W. 2013. Multiple nuclear and mitochondrial DNA sequences provide new insights into the phylogeny of South African Lacertids (Lacertidae, Eremiadinae). Journal of Zoological Systematics and Evolutionary Research, 51: 132–143. doi: 10.1111/jzs.12012 - get paper here
  • FitzSimons, V.F. 1943. The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp.
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  • Pietersen, Darren W. Pietersen, Errol W. Haacke, Wulf D. 2013. First herpetological appraisal of the Parque Nacional de Banhine, Gaza Province, southern Mozambique. Annals of the Ditsong National Museum of Natural History 3: 153-163 - get paper here
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  • Schmidt, A. D. 2005. Habitatnutzung, Raumbedarf und Mobilität von Ichnotropis squamulosa PETERS 1854 (Reptilia, Lacertidae) in der Trockensavanne Südafrikas. Sauria 27 (4): 3-14 - get paper here
  • SCHMIDT, A.D. 2006. Zur Populationsökologie von Ichnotropis squamulosa PETERS, 1854 (Reptilia, Lacertidae) in der Trockensavanne Südafrikas. Sauria 28 (1): 31-38 - get paper here
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