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Phrynocephalus reticulatus EICHWALD, 1831

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
SubspeciesPhrynocephalus reticulatus bannikovi DAREVSKY, RUSTAMOV & SHAMMAKOV 1976
Phrynocephalus reticulatus reticulatus EICHWALD 1831 
Common NamesE: Reticulated toad-headed agama 
SynonymAgama ocellata LICHTENSTEIN 1823: 143
Phrynocephalus reticulatus EICHWALD 1831: 186
Phrynocephalus reticulatus — DUMÉRIL & BIBRON 1837: 515
? Phrynocephalus ocellatus — DUMÉRIL & BIBRON 1837: 516
Phrynocephalus caudivolvulus reticulatus
Phrynocephalus nikolskii BEDRIAGA in NIKOLSKY 1905: 463
Phrynocephalus boulengeri — BEDRIAGA in NIKOLSKY 1905: 466
Phrynocephalus nikolskii BEDRIAGA 1907: 210
Phrynocephalus boulengeri BEDRIAGA 1907: 213
Phrynocephalus caudivolvulus var. reticulatus — BEDRIAGA 1909: 367
Phrynocephalus reticulatus — SMITH 1935: 231
Phrynocephalus ocellatus — DUNAYEV 1996
Phrynocephalus reticulatus — BARTS & WILMS 2003
Phrynocephalus ocellatus — SZCZERBAK 2003
Phrynocephalus (Phrynocephalus) reticulatus reticulatus — BARABANOV & ANANJEVA 2007
Phrynocephalus ocellatus — SINDACO & JEREMČENKO 2008

Phrynocephalus reticulatus bannikovi DAREVSKY; RUSTAMOV & SHAMMAKOV 1976
Phrynocephalus bannikovi — LICHNOVA 1992: 7
Phrynocephalus (Phrynocephalus) reticulatus bannikovi — BARABANOV & ANANJEVA 2007
Phrynocephalus reticulatus bannikovi — SINDACO & JEREMČENKO 2008
Phrynocephalus ocellatus bannikovi — SHESTOPAL & SCHCHERBINA 2024 
DistributionS Kazakhstan ?, W Uzbekistan, Turkmenistan, N India (Jammu and Kashmir)

Type locality: Valley of the River Amudarya and East Coast of the Caspian Sea

bannikovi: NW Turkmenistan (Krasnovodsky Peninsula); Terra typica: “Tuarkyr Mountains, NW Turkmenistan”.  
Reproductionoviparous 
TypesHolotype: ZISP (ZIL) 8720, Banks of the Oxus.
Holotype: ZISP 18505, male, “Tuarkyr mountains, north-western Turkmenia” [Tuarkyr mountains, Balkan province, Turkmenistan]. Leg: Ch. Atajev and S. Sham- makov, 23.IX.1971 [bannikovi]
Lectotype: ZMB 787, "Kleine Tartarei", leg. Eversmann Paralectotypus: ZMB 788, "Bucharei", leg. Eversmann; Terra typica: Tartarei [ocellata] 
Diagnosis 
CommentSynonymy partly after KHALIKOV & ANANJEVA (pers. comm.). Not listed as such by DAS 1996. BOULENGER 1885 synonymized Agama ocellata LICHTENSTEIN 1823 with P. guttatus. GOLUBEV (1991) stated that this name is the senior synonym for Phrynocephalus reticulatus Eichwald, 1831. Therefore he placed the junior name in the synonymy of the resurrected Phrynocephalus ocellatus (Lichtenstein In Eversmann, 1823). However, the name reticulatus has now been consistently used for more than one century by many authors, and should be protected (BARABANOV & ANANJEVA 2007).

Distribution: SMITH (1935) questions the record from Ladakh. SINDACO & JEREMCENKO 2008 also state that records from the Fergana Valley and from Kashmir are erroneous. Not in Afghanistan fide Wagner et al. 2017.

NCBI taxon ID: 1585972 [bannikovi] 
References
  • Ananjeva, Natalia B.;Tuniyev, Boris S. 1992. Historical biogeography of the Phrynocephalus species of the USSR. Asiatic Herpetological Research 4: 76-98 - get paper here
  • BARABANOV, A.V. & N.B. ANANJEVA 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa 1399: 1-56 - get paper here
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Bedriaga, J.V. 1909. Amphibien und Reptilien. In: Wissenschaftliche Resultate der Reisen N.M. Przewalskijs durch Zentralasien. Zoologische Teil. Band 3. Part 1. Lacertilia. (in Russian). Sankt-Petersburg, Vol 3 (1) 3: 73-102, 367 - get paper here
  • Bedriaga, JACQUES VON 1907. Neue Saurier aus Russisch-Asien. Annu. Mus. Zool. Acad. Impér. Sci. St.-Petersbourg, 10 [1905]: 210-243 - get paper here
  • Darevsky, I.S., Rustamov, A.K. & Shammakov, S. 1976. Status and distribution of Phrynocephalus reticulatus Eichwald (Sauria, Agamidae) in Central Asia. [in Russian] Teoreticheskije i prikladnyije aspekty okhrany prirody i okhotovedenija, Moscow, 84, 113–119
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Dunayev, E. A. 1996. On the possible use of the Ethological features in the taxonomy and phylogeny of toad agamas, Phrynocephalus (Reptilia, Agamidae). Russ. J. Herpetol. 3 (1): 32-38 - get paper here
  • Eichwald, E. 1831. Zoologia specialis, quam expositis animalibus tum vivis, tum fossilibus potissimuni rossiae in universum, et poloniae in specie, in usum lectionum publicarum in Universitate Caesarea Vilnensi. Zawadski, Vilnae. - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Hoogmoed, M.S. 1980. Herpetologische waarnemingen in Ghana [part 3]. Lacerta 38 (5): 42-48 - get paper here
  • Milto, Konstantin D.; Andrei V. Barabanov 2012. A Catalogue of the Agamid and Chamaeleonid Types in the Collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ. J. Herpetol. 19 (2): 155-170 - get paper here
  • Nikolsky, A. M. 1905. Herpetologia rossica. Mem. Acad. Imp. Sci., Phys.-Math. Classe, St. Pétersbourg, XVII (1): 1-518
  • Sahi D.N., Koul S. 2020. Annotated List of Amphibians and Reptiles of Jammu and Kashmir State. In: Dar G. & Khuroo A. (eds) Biodiversity of the Himalaya: Jammu and Kashmir State. Topics in Biodiversity and Conservation, vol 18. Springer, Singapore - get paper here
  • Shestopal A. A., Shcherbina A. A. 2024. Distribution of reptiles in Northwestern Turkmenistan. [in Russian] Current Studies in Herpetology 24 (1-2): 102 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Waltner, R.C. 1975. Geographical and altitudinal distribution of amphibians and reptiles in the Himalayas. Cheetal (Dehra Dun, India) 16(1): 17-25; 16(2): 28-36; 16(3): 14-19; 16(4): 12-17.
 
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