Phymaturus williamsi LOBO, LASPIUR & ACOSTA, 2013

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Higher Taxa	Liolaemidae, Iguania, Sauria, Squamata (lizards)
Subspecies
Common Names
Synonym	Phymaturus williamsi LOBO, LASPIUR & ACOSTA 2013
Distribution	NW Argentina Type locality: Quebrada vallecito, 40 km W of Calingasta town, Calingasta department, San Juan province. 31°11’21’’ S; 69°42’15.1’’ W, 3000 m elevation.
Reproduction	viviparous (not imputed, fide Zimin et al. 2022)
Types	Holotype: MCN-UNSa (=MCN) 2820. Male. A. Laspiur, R. Acosta & J. C. Acosta cols. 11/05/2008. Paratypes: MCN 2808-10, 2812-14, 2815, 2816-17, 2821. Same data as holotype. MCN 3259-65. Quebrada Vallecito, 40 km W of Calingasta town, Calingasta department, San Juan province. 31o11’21’’S; 69o42’15.1’’W, 3000m asl. A. Laspiur & J. C. Acosta cols.
Diagnosis	Diagnosis: Phymaturus williamsi sp. nov. differs from all other members of the Puna clade because: exhibits an “aggregate” dorsal pattern, unlike the ho- mogeneous spray of most Puna species, lacks enlarged scales on the anterior margin of the antehumeral fold and in the centre of chest, flank coloration in females is absent, females of Phymaturus williamsi sp. nov. lack white transversal stripes on the dorsal pattern, preocular scale in contact with canthal scale in Phymaturus williamsi sp. nov., rostral scale can be divided in Phymaturus williamsi sp. nov. and shows the largest number of scales counted around midbody within the Puna subclade (x= 213.4; 186-235).
Comment	Abundance: only known from its original description (Meiri et al. 2017).
Etymology	We name this new species in honour of our Argentine colleague and friend Jorge Williams, in recognition of his effort and dedication for the development of herpetology in our country.
References	Castro, Susana A.; Jorgelina M. Boretto, Graciela M. Blanco, and Juan C. Acosta 2018. Adjustment of the Reproductive Activity of Vulnerable Lizard Phymaturus williamsi at High Altitudes. Herp. Cons. Biol. 13 (1): 283–293. - get paper here Erostarbe, A. V. and Acosta, J. C. 2020. Phymaturus williamsi. Parturition behavior. Herpetological Review 51: 131 - get paper here Fava, G A; Acosta, J C; Victorica, A E; Martínez, T & Rodríguez, M 2018. Phymaturus williamsi (Predator-prey interaction). Herpetological Review 49 (2): 331-332 - get paper here LASPIUR, A., MEDINA, M., AUSAS, M. S., ACOSTA, J. C., KRENZ, J. D., & IBARGÜENGOYTÍA, N. R. 2024. Thermal niches and activity periods in syntopic Phymaturus and Liolaemus lizards from the Andes, Argentina. Anais da Academia Brasileira de Ciências, 96, e20191190 - get paper here LOBO, FERNANDO; ALEJANDRO LASPIUR, JUAN CARLOS ACOSTA 2013. Description of new andean species of the genus Phymaturus (Iguania: Liolaemidae) from Northwestern Argentina. Zootaxa 3683: 117–132 - get paper here Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here Troncoso-Palacios, Jaime and Damien Esquerré 2014. A new species of Phymaturus of the P. mallimaccii Group from the Andes of central Chile (Iguania: Liolaemidae). Phyllomedusa 13 (1):3–15 - get paper here Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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