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Ptyas carinata (GÜNTHER, 1864)

IUCN Red List - Ptyas carinata - Least Concern, LC
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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Keeled Rat Snake
G: Gekielte Rattenschlange
Chinese: 黑网乌梢蛇 
SynonymCoryphodon carinatus GÜNTHER 1858: 112 (invalid)
Zaocys carinatus GÜNTHER 1864
Zaocys (Zapyrus) carinatus — BOETTGER 1887
Zaocys tenasserimensis SCLATER 1891: 238
Zaocys carinatus — BOULENGER 1893: 377
Ptyas carinatus — TAYLOR 1922: 136
Ptyas carinatus — WALL 1923
Zaocys carinatus — SMITH 1943
Ptyas carinatus — TAYLOR 1965: 736
Zaocys carinatus — GRANDISON 1978: 291
Ptyas carinatus — DAVID & VOGEL 1996
Zaocys carinatus — MANTHEY & GROSSMANN 1997: 402
Ptyas carinatus — COX et al. 1998: 55
Zaocys carinatus — GRISMER et al. 2002
Zaocys carinatus — ROOIJEN & ROOIJEN 2002
Ptyas carinatus — MALKMUS et al. 2002
Ptyas carinata — DAVID & DAS 2004
Zaocys carinatus — ZHAO 2006
Ptyas carinata — WALLACH et al. 2014: 615 
DistributionBurma (Myanmar), China (Yunnan),
Indonesia (Sumatra, Java, Borneo), Laos (Oudomxay, Bokèo, Xaignabouli),
Malaysia (Malaya and East Malaysia, Pulau Tioman),
Philippines (Palawan), Singapore Island,
Thailand, Vietnam, Cambodia

Type locality: Borneo  
Reproductionoviparous 
TypesHolotype: BMNH (presumed, status unclear fide P. Campbell, pers. comm., March 2019). 
DiagnosisDiagnosis: Maxillary teeth, 22-26; internasals two thirds of the length of prefrontals; two to four loreals; eight to ten supralabials, fourth and fifth or fifth and sixth bordering orbit. Scale-formula: 18, 16 (14), 12; four to six median scalerows keeled. (Taylor 1965: 736)


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CommentSynonymy: The description of this species has constantly been credited to Günther (1858) but as shown by Dubois & David (2020), the nomen created by this author, although widely used, is not valid as it is an invalid objective junior synonym of Coluber dhumnades Cantor, 1842.

Habitat: partly arboreal (Harrington et al. 2018).

This is the largest Asian colubrid snake as its maximum total length is 399 centimeters (Das 2012). 
EtymologyNamed after Latin “carinatus” = keeled (like a ship). 
References
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  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
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  • Das, Indraneil; Jye Wen Wong 2021. Predation on Gonocephalus liogaster (Agamidae) by Ptyas carinata (Colubridae) in Sarawak, Borneo. Herpetology Notes 14: 349-351 - get paper here
  • David, P. & I. Das 2004. On the grammar of the gender of Ptyas Fitzinger, 1843 (Serpentes: Colubridae). Hamadryad 28 (1&2): 113-116 - get paper here
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