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Sepsiscus pluto (INGRAM, 1977)

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Cape York Worm-skink 
SynonymAnomalopus (Vermiseps) pluto INGRAM 1977
Anomalopus pluto — GREER & COGGER 1985: 24
Coeranoscincus pluto — WELLS & WELLINGTON 1985: 26
Anomalopus pluto — COGGER 2000: 385
Anomalopus pluto — WILSON & SWAN 2010
Sepsiscus pluto — HUTCHINSON et al. 2021
Suppressascincus pluto 
DistributionAustralia (Queensland)

Type locality: McDonald Crossing, Cockatoo Creek, 115 km S of Bamaga, Cape York, in 11° 33’ S, 142° 26’ E, Qld.  
Reproductionoviparous 
TypesHolotype: QM J26261 
DiagnosisDiagnosis (genus). Body serpentiform; limbless. Eyelids moveable, scaly; ear opening absent but external ear differentiated as a vertically elongate trough with small weakly ossified scales (Fig. 3C in Hutchinson et al. 2021).
Head shields (Fig. 4C); nasals separated medially, nasal fused to first supralabial; prefrontals present, small, widely separated; loreals two; supraoculars two; supraciliaries three (not in a continuous series) with the first two separated from the third by the second supraocular projecting laterally into the supraciliary row; parietal margined laterally and posteriorly by a single upper secondary temporal in contact with a single enlarged nuchal; subocular scale row much reduced; frontoparietals separate; postmental contacting a single infralabial on each side.
Parietal and pulmonary peritoneum not pigmented; right oviduct absent (Greer and Cogger, 1985). 
CommentLimb morphology: 0 digits 0 toes (Limbless, Singhal et al. 2018, Cogger 2014)

Type species: Anomalopus (Vermiseps) pluto INGRAM 1977 is the type species of the genus Sepsiscus HUTCHINSON et al. 2021.

Morphology: Hutchinson et al. 2021 present a table of morphological character states across 20 Australian sphenomorphine skinks, including this genus. 
EtymologyNamed after the god of the underworld, Pluto.

The genus was named after Latin seps, classically, a small but dangerous snake, but in the herpetological literature associated with limb-reduced skinks, and -iscus, a diminutive, thus ‘‘little seps.’’ 
References
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Couper, P., Covacevich, J., Amey, A. & Baker, A. 2006. The genera of skinks (Family Scincidae) of Australia and its island territories: diversity, distribution and identification. in: Merrick, J.R., Archer, M., Hickey, G.M. & Lee, M.S.Y. (eds.). Evolution and Zoogeography of Australasian Vertebrates. Australian Scientific Publishing, Sydney, pp. 367-384
  • Greer A E; Cogger H G 1985. Systematics of the reduce-limbed and limbless skinks currently assigned to the genus Anomalopus (Lacertilia: Scincidae). Rec. Austral. Mus. 37(1) 1985: 11-54 - get paper here
  • Hutchinson, M. N., Couper, P., Amey, A., & Wilmer, J. W. 2021. Diversity and Systematics of Limbless Skinks (Anomalopus) from Eastern Australia and the Skeletal Changes that Accompany the Substrate Swimming Body Form. Journal of Herpetology 55 (4): 361-384 - get paper here
  • Ingram G J 1977. A new species of legless skink Anomalopus pluto from Cape York Peninsula, Queensland. Victorian Naturalist 94 (2): 52-53 - get paper here
  • Singhal, Sonal; Huateng Huang, Maggie R. Grundler, María R. Marchán-Rivadeneira, Iris Holmes, Pascal O. Title, Stephen C. Donnellan, and Daniel L. Rabosky 2018. Does Population Structure Predict the Rate of Speciation? A Comparative Test across Australia’s Most Diverse Vertebrate Radiation. The American Naturalist - get paper here
  • Skinner, Adam; Mark N. Hutchinson, Michael S.Y. Lee 2013. Phylogeny and Divergence Times of Australian Sphenomorphus Group Skinks (Scincidae, Squamata). Molecular Phylogenetics and Evolution 69 (3): 906–918 - get paper here
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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