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Tytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL & QUAH, 2018

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Sungai Bubu Forest Skink
Malay: Mengkarung Hutan Sungai Bubu 
SynonymTytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL4& QUAH 2018 
DistributionPeninsular Malaysia (Terengganu)

Type locality: hilly area near Sungai Bubu, Sekayu, Hulu Terengganu, Terengganu State, Peninsular Malaysia (4.9710° N, 102.9531° E; 174 m in elevation  
Reproduction 
TypesHolotype: LSUHC 13858, Adult male, collected by M. Afiq-Shuhaimi and Baizul-Hafsyam B. S. on 5 August 2017. 
DiagnosisDiagnosis Tytthoscincus monticolus sp. nov. can be differentiated from all other species of Tytthoscincus in the upland clade by having the combination of 8`1, superciliaries; a deeply set, unpigmented tympanum; enlarged pectoral scales; 31 midbody scale rows; 69 paravertebral scales; 68 ventral scales; keeled, subdigital lamellae; seven subdigital lamellae on the third finger; 12 subdigital lamellae on the fourth toe; EO/HL = 0.16; HL/SVL = 0.17; AXG/SVL = 0.55; FL/SVL = 0.22; HDL/SVL = 0.33; and a maximum SVL of 36.6 mm (Tables 5,6). All characters are scored across all other Tytthoscincus and species of Sphenomorphus suspected of being Tytthoscincus in Grismer et al. (2016a:237) and Karin et al. (2016:416).


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CommentHabitat: leaf-litter in hilly areas along riparian systems in lowland dipterocarp forest 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here
  • GRISMER, L. LEE; PERRY L. JR. WOOD, AMIRRUDIN B. AHMAD, B. S. BAIZUL-HAFSYAM, M. AFIQ-SHUHAIMI, SYED A. RIZAL, EVAN S. H. QUAH 2018. Two new Tytthoscincus Linkem, Diesmos, & Brown (Squamata; Scincidae) from Peninsular Malaysia and another case of microsyntopy between ecologically specialized, unrelated, leaf-litter species. Zootaxa 4425 (1): 087–107 - get paper here
  • KARIN, BENJAMIN R.; INDRANEIL DAS, AARON M. BAUER 2016. Two new species of diminutive leaf-litter skinks (Squamata: Scincidae: Tytthoscincus) from Gunung Penrissen, Sarawak, Malaysia (northern Borneo). Zootaxa 4093 (3): 407–423 - get paper here
  • Syafiq, Muhamad Fatihah; Baizul Hafsyam Badli-Sham, Noor-Shahirah Ibrahim, Lukman H. B. Ismail, Muhammad Afif Mohd Amin, Goh Shu Xian, Rabiatul ‘Aqilah Mohamad Ariffin, Muhammad Afiq-Suhaimi, Lok Kar Men, Dila Adilina Danelo, Mohamad Aqmal-Naser, Muh 2024. TAXONOMIC COMPOSITION, DIVERSITY, AND CONSERVATION STATUS OF REPTILIAN FAUNA AT SEKAYU LOWLAND FORESTS, TERENGGANU, PENINSULAR MALAYSIA. Russian Journal of Herpetology Vol. 31, No. 1, 2024, pp. 14 – 23 - get paper here
 
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