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Varanus niloticus (LINNAEUS, 1766)

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Higher TaxaVaranidae, Platynota, Varanoidea, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Nile Monitor, Water Leguaan
G: Nilwaran 
SynonymLacerta monitor LINNAEUS 1758: 201 (nomen rejectum)
Lacerta nilotica — LINNAEUS 1766: 369
Stellio saurus LAURENTI 1768: 56
Lacerta capensis SPARRMAN 1783: 749
Lacertus tupinambis LACÉPÈDE 1788 (in part)
Lacerta nilotica — MEYER 1795: 29
Tupinambis elegans DAUDIN 1802 (in part)
Tupinambis stellatus DAUDIN 1802 (in part)
Tupinambis ornatus DAUDIN 1803: 353
Monitor niloticus — LICHTENSTEIN 1818
Monitor pulcher LEACH in BOWDICH 1819
Varanus niloticus — FITZINGER 1826: 50
Varanus niloticus — DUMÉRIL & BIBRON 1836: 476
Monitor elegans senegalensis — SCHLEGEL 1839: X
Monitor elegans capensis — SCHLEGEL 1839: 76
Monitor elegans senegalensis — SCHLEGEL 1844
Varanus ornatus — GRAY 1845
Varanus niloticus — BOULENGER 1885: 317
Varanus niloticus — SCHMIDT 1919: 485
Varanus (Polydaedalus) niloticus — MERTENS 1942
Varanus (Polydaelus) niloticus ornatus — MERTENS 1942
Varanus niloticus — AUERBACH 1987: 136
Varanus niloticus — DE LISLE 1996: 131
Varanus ornatus — BÖHME & ZIEGLER 1997
Varanus (Polydaedalus) ornatus — SCHMITZ et al. 2000
Varanus niloticus — AST 2001
Varanus ornatus — PAUWELS et al. 2004
Varanus niloticus — TRAPE, CHIRIO & TRAPE 2012
Varanus (Polydaedalus) ornatus — BUCKLITSCH et al. 2016: 50
Varanus (Polydaedalus) niloticus — BUCKLITSCH et al. 2016: 50
Varanus ornatus — PAUWELS et al. 2016
Varanus niloticus — SPAWLS et al. 2018: 309
Varanus ornatus — PAUWELS et al. 2019
Varanus ornatus — BÖHME 2023
Varanus ornatus — BÖHME et al. 2023 
DistributionRepublic of South Africa (Eastern Cape etc.), Swaziland, Namibia, Botswana, Tanzania, Mozambique, Zimbabwe, Zambia, Angola, Malawi, Tanzania, Gabon, W/N/S Democratic Republic of the Congo (Zaire), Kenya, Uganda, Cameroon, Central African Republic, Ethiopia, Eritrea, Somalia, Sudan (Jumhūriyyat), Republic of South Sudan (RSS), Chad, Egypt, Liberia, Ivory Coast, Ghana, Togo, Benin, Burkina Faso, Niger, Nigeria, Mali, Mauritania, Senegal, Gambia (HÅKANSSON 1981), Guinea (Conakry), Equatorial Guinea

ornatus: Guinea, Togo, Benin, Liberia, Sierra Leone, Guinea-Bissau, Democratic Republic of the Congo (Zaire; except in the S), Cameroon, Gabon (BLANC & FRETEY 2000), Ivory Coast, Central African Republic; Type locality: Malimbe, Cameroon.

Introduced to USA (Florida)

Type locality: “Indiis” (Linnaeus 1758); Type locality: “in Ægypto” [= Egypt] (fide Hasselquist 1757, cited in Marques et al. 2018: 277).  
Reproductionmaybe parthenogenetic (Hennessy 2010). 
TypesSyntype: NRM 9185; Specimen mentioned by Hasselquist (1757) (collector F. Hasselquist), presumed lost fide Marques et al. 2018.
Varanus niloticus: Bocage
Syntypes: MNHN [Tupinambis stellatus]
Holotype: MNHN 7471 [ornatus] 
DiagnosisDiagnosis: (1) Large lizard up to approximately 80 cm SVL and about 200 cm total length. (2) Tail laterally compressed with a low dorsal crest. (3)
A total of 136 to 183 scales around the midbody. (4) Basic dorsal color of adults gray brown to olive-brown with light yellow ocelli and bands on head, back, limbs, and tail. Belly and throat are paler, with black bars. (5) V. niloticus is characterized by six to nine crossbands or rows of yellow ocelli on the back between foreand hind limbs, whereas V. ornatus normally has only five bands. (Lenz in Pianka et al. 2004)


Additional details (169 characters) are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy: The status of V. ornatus remains somewhat contentious. Böhme and Ziegler (1997) elevated V. ornatus to full species level based on differing pigmentation patterns, tongue coloration, genital morphology, and slight differences in scale counts. However, Dowell et al. (2015) synonymized V. ornatus with V. niloticus as both are genetically indistinguishable and because morphological characters largely overlap. However, Dowell et al. found distinct clades within V. niloticus across its huge range. SCHMIDT’s Varanus niloticus might represent both V. niloticus (specimens from savanna localities) and V. ornatus (specimens from rain forest localities). ICZN opinion 540 rejected Lacerta monitor LINNAEUS 1758.

Distribution: For a detailed distribution map and locality data see BAYLESS (2002). The monitor in Thieme 2004: 46 is not V. ornatus but rather V. niloticus. For a map with localities in Equatorial Guinea see SÁNCHEZ-VIALAS et al. 2022.

Conservation: although not threatened in the whild due to its large range, this species has been driven to local extinction by overexploitation for skins (Hinsley et al. 2023). 
References
  • Abdelkarim, A. A., Kanno, I. O., & Musa, F. I. 2024. Observation on identification of birds and reptile in Taya-Basonda–Al galabat National Park, Gedaref state-Sudan. Journal for Nature Conservation, 126614 - get paper here
  • Abdelkarim, A.A., Kanno, I.O. & Musa, F.I. 2024. Observation on identification of birds and reptile in Taya - Basonda – Al galabat National Park, Gedaref State - Sudan. Journal for Nature Conservation 79, 126614 - get paper here
  • Amer, Sayed A. M. and Yoshinori Kumazawa 2008. Timing of a mtDNA gene rearrangement and intercontinental dispersal of varanid lizards. Genes Genet. Syst. 83: 275–280 - get paper here
  • Ast, Jennifer C. 2001. Mitochondrial DNA Evidence and Evolution in Varanoidea (Squamata). Cladistics 17 (3): 211-226 [erratum in 18 (1):125] - get paper here
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Baha el Din, S. 2006. A guide to the reptiles and amphibians of Egypt. The American University in Cairo Press, Cairo. xvi + 359 pp.
  • Barnes, Keith 2016. Animals of Kruger National Park. Princeton University Press, ca. 180 pp. - get paper here
  • Barnett, Linda K. 2001. The herpetofauna of Abuko Nature Reserve, the Gambia. Herpetological Bulletin (77): 5-1 - get paper here
  • Barnett, Linda K. & Emms, Craig 2005. Common reptiles of The Gambia. Rare Repro, Hailsham, East Sussex, 24 pp.
  • BATES, M.F. 2010. Varanus niloticus (Linnaeus, 1766). African Herp News (52): 25-26 - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001]
  • Bayless, Mark K. 2002. Monitor lizards: a pan-African check-list of their zoogeography (Sauria: Varanidae: Polydaedalus). Journal of Biogeography 29: 1643–1701 - get paper here
  • Behangana, M.; C. Aluziyo, D dendi, L. Luiselli, D. F. Hughes, E. Greenbaum 2019. HERPETOFAUNAL DIVERSITY AND COMMUNITY STRUCTURE AT LAKE NABUGABO AND SURROUNDING RAMSAR AREA WETLANDS IN SOUTHERN UGANDA. Vie et milieu - life and environment 69 (2-3): 1-N - get paper here
  • Behangana, Mathias; Richard Magala, Raymond Katumba, David Ochanda, Stephen Kigoolo, Samuel Mutebi, Daniele Dendi,, Luca Luiselli, and Daniel F. Hughes 2020. Herpetofaunal diversity and community structure in the Murchison Falls-Albert Delta Ramsar site, Uganda: Herpetofaunal diversity. European Journal of Ecology, 6(2)
  • Bennett, D. 2002. Diet of Juvenile Varanus niloticus (Sauria: Varanidae) on the Black Volta River in Ghana Journal of Herpetology 36 (1): 116-117. - get paper here
  • Bennett, Daniel 1998. Monitor Lizards Natural History, Biology and Husbandry. Edition Chimaira, Frankfurt - get paper here
  • Blanc, P. & Fretey, T. 2000. Les reptiles de la Réserve de La Lopé et de la Foret des Abeilles (Gabon). Bull. Soc. Zool. France 125 (4): 281-292
  • Böhme, W. & T. ZIEGLER 1997. A taxonomic review of the Varanus (Polydaedalus) niloticus (Linnaeus, 1766) species complex. The Herpetological Journal 7: 155-162. - get paper here
  • Böhme, Wolfgang 2023. Der Bonner afrotropische Regenwald und seine herpetologischen Bewohner. Elaphe 2023 (5): 56-61
  • Böhme, Wolfgang, Mark-Oliver Rödel, Christian Brede & Philipp Wagner 2011. The reptiles (Testudines, Squamata, Crocodylia) of the forested southeast of the Republic Guinea (Guinée forestière), with a country-wide checklist. Bonn zoological Bulletin 60 (1): 35-61 - get paper here
  • Böhme, Wolfgang; Eberhard Fischer & Harald Hinkel 2023. Reptilien aus dem Irangi Forest, Provinz Kivu-Süd, Demokratische Republik Kongo. Sauria 45 (4): 29
  • Böhme,W. 2003. Checklist of the living monitor lizards of the world (family Varanidae). Zoologische Verhandelingen 341: 6-43 - get paper here
  • Bonetti, Mathilde 2002. 100 Sauri. Mondadori (Milano), 192 pp. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Bourquin, O. 1991. Life History Note: Varanus niloticus: Reproduction. J. Herp. Assoc. Africa (39): 21-21 - get paper here
  • Branch, W. R. 1982. Hemipeneal Morphology of Platynotan Lizards. Journal of Herpetology 16 (1): 16-38 - get paper here
  • Branch, W.R. 1992. Life History Notes - Varanus niloticus niloticus. J. Herp. Assoc. Africa (41): 40-40 - get paper here
  • Branch, W.R. & Rödel, M.-O. 2003. Herpetological survey of the Haute Dodo and Cavally forests, western ivory Coast, Part II: Trapping, results and reptiles. Salamandra 39 (1): 21-38 - get paper here
  • Branch, W.R.; Erasmus, H. 1982. Notes on Reproduction in South African Water Monitor, Varanus niloticus niloticus. J. Herp. Assoc. Africa (28): 4-7 - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Broadley, D.G. 1962. On some reptile collections from the North-Western and North-Eastern Districts of Southern Rhodesia 1958-1961, with descriptions of four new lizards. Occ. Pap. Nat. Mus. South. Rhodesia 26 (B): 787-843
  • Broadley, D.G. 1991. The Herpetofauna of Northern Mwinilunga Distr., Northw. Zambia. Arnoldia Zimbabwe 9 (37): 519-538
  • Broadley, Donald G. and F. P. D. Cotterill. 2004. The reptiles of southeast Katanga, an overlooked 'hot spot'. [Congo]. African Journal of Herpetology 53 (1): 35-61. - get paper here
  • Brygoo, E. R. 1987. Les types de Varanides (Reptiles, Sauriens) du Museum national d'Histoire naturelle. Catalogue critique. Bulletin du Museum national d'Histoire naturelle, Paris, 4e ser., 9, sect. A (Zool., BioI. & Ecol. anim.), (2), supplement, 21-38.
  • BUCKLITSCH, YANNICK; WOLFGANG BÖHME, ANDRÉ KOCH 2016. Scale Morphology and Micro-Structure of Monitor Lizards (Squamata: Varanidae: Varanus spp.) and their Allies: Implications for Systematics, Ecology, and Conservation. . Zootaxa 4153 (1): 001–192 - get paper here
  • Buffrénil, V. de & J. Castanet 2000. Age Estimation by Skeletochronology in the Nile Monitor (Varanus niloticus), a Highly Exploited Species Journal of Herpetology 34 (3): 414-424. - get paper here
  • Buffrénil, Vivian de and Georges Hémery 2002. Variation in longevity, growth, and morphology in exploited Nile Monitors (Varanus niloticus) from Sahelian Africa. Journal of Herpetology 36 (3): 419-426 - get paper here
  • Buruwate, T. C., & Lloyd-Jones, D. J. 2024. Amphibian and Reptile Diversity of Niassa Special Reserve, Northern Mozambique. Journal of East African Natural History, 113(1), 1-18 - get paper here
  • Carlino, P. & Pauwels, O.S.G. 2015. An Updated Reptile List of Ivindo National Park, the Herpetofaunal Hotspot of Gabon. Bull. Chicago Herp. Soc. 50(3): 25-39 - get paper here
  • Ceríaco, Luis M. P., Aaron M. Bauer, David C. Blackburn and Ana C. F. C. Lavres. 2014. The herpetofauna of the Capanda Dam Region, Malanje, Angola. Herpetological Review 45 (4): 667-674 - get paper here
  • Charlton, P. E. 1973. Observations on the Nile Monitor Lizard, Varanus niloticus in Natal. J. Herp. Assoc. Africa (11): 19-19 - get paper here
  • Chirio, L. 2009. Inventaire des reptiles de la région de la Réserve de Biosphère Transfrontalière du W (Niger/Bénin/Burkina Faso: Afrique de l’Ouest). [Herpetological survey of the W Transfrontier Biosphere Reserve area (Niger/Benin/Burkina Faso: West Africa]. Bull. Soc. Herp. France (132): 13-41 - get paper here
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Chirio, Laurent and Ivan Ineich 2006. Biogeography of the reptiles of the Central African Republic. African Journal of Herpetology 55(1):23-59. - get paper here
  • Cimatti, E. 2005. Scaly encounters in Kenya. Reptilia (GB) (42): 65-69 - get paper here
  • Conradie W and Branch WR. 2016. The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation 10 (2) [Special Section]: 6–36 - get paper here
  • Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J, Júlio T, Neef G. 2022. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles. Amphibian & Reptile Conservation 16(2): 181–214 (e322) - get paper here
  • CONRADIE, W., DOUCETTE-RIISE, S., VANHOOYDONCK, B., ENGELBRECHT, H., MEASEY, G.J., & TOLLEY, K. 2011. Herpetological survey of Rooipoort Nature Reserve, Northern Cape, South Africa. African Herp News (53): 35-41 - get paper here
  • CONRADIE, W., ENGELBRECHT, H., HERREL, A., MEASEY, G.J., NIELSEN, S.V., V ANHOOYDONCK, B., & TOLLEY, K.A. 2011. Reptile survey of Venetia Limpopo Nature Reserve, Limpopo Province - South Africa. African Herp News (55): 6-12 - get paper here
  • Conradie, Werner; Brian Reeves , Neil Brown, & Jan A. Venter 2016. Herpetofauna of the Oviston, Commando Drift and Tsolwana nature reserves in the arid interior of the Eastern Cape Province, South Africa. Indago 32: 81-98 - get paper here
  • Dalhuijsen, Kim; William R. Branch, and Graham J. Alexander 2014. A Comparative Analysis of the Diets of Varanus albigularis and Varanus niloticus in South Africa. African Zoology Apr 2014, Vol. 49, No. 1: 83-93. - get paper here
  • Daudin, F. M. 1802. Histoire Naturelle, génerale et particulièredes reptiles, ouvrage faisant suite, a l'histoiure naturelle, générale et particulière composée par LECLERC DE BUFFON, et redigée par C. S. SONNINI, vol. 3. F. Dufart, Paris, 452 pp. - get paper here
  • Daudin, F. M. 1803. Histoire Naturelle, Générale et Particulière des Reptiles, Vol. 8. F. Dufart, Paris, 439 pp. - get paper here
  • Dowell, Stephanie A.; Daniel M. Portik, Vivian de Buffrénil, Ivan Ineich, Eli Greenbaum, Sergios-Orestis Kolokotronis, Evon R. Hekkala 2015. Molecular data from contemporary and historical collections reveal a complex story of cryptic diversification in the Varanus (Polydaedalus) niloticus Species Group. Molecular Phylogenetics and Evolution 94: 591–604 - get paper here
  • Duméril, A.M. C. and G. Bibron. 1836. Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 3. Libr. Encyclopédique Roret, Paris, 528 pp. - get paper here
  • Earley, Ryan L.; Omar Attum, Perri Eason 2002. Varanid combat: perspectives from game theory. Amphibia-Reptilia 23 (4): 469-485 - get paper here
  • Ernst R, Lautenschläger T, Branquima MF, Hölting M 2020. At the edge of extinction: a first herpetological assessment of the proposed Serra do Pingano Rainforest National Park in Uíge Province, northern Angola. Zoosystematics and Evolution 96(1): 237-262 - get paper here
  • Fitzinger, L. 1826. Neue Classification der Reptilien nach ihren natürlichen Verwandtschaften nebst einer Verwandschafts-Tafel und einem Verzeichnisse der Reptilien-Sammlung des K. K. Zoologischen Museums zu Wien. J. G. Heubner, Wien, 66 pp. - get paper here
  • FRÝDLOVÁ, PETRA and DANIEL FRYNTA 2010. A test of Rensch’s rule in varanid lizards. Biological Journal of the Linnean Society 100: 293–306 - get paper here
  • Gibbons, Whit; Judy Greene, and Tony Mills 2009. LIZARDS AND CROCODILIANS OF THE SOUTHEAST. University of Georgia Press, 240 pp.
  • Gonwouo, Nono LeGrand; Tchassem Fonkwa Arnaud Marius, Doherty-Bone Thomas, Rödel Mark-Oliver 2021. Amphibian and reptiles of a proposed iron ore mining concession in southern Cameroon. Herpetology Notes 14: 1051-1065
  • Gossmann, V.; Lötters, S.; Obame, F. & Böhme, W. 2002. Zur Herpetofauna Gabuns. Teil II: Kommentierte Artenliste der gefundenen Reptilien, Bemerkungen zur Artenvielfalt. Herpetofauna 24 (136): 19-33 - get paper here
  • Grützner, T. 2003. Nachzucht von Varanus ornatus (DAUDIN 1803). Elaphe 11 (4): 25-34
  • Günther, A. 1861. On the anatomy of Monitor niloticus from Western Africa, and of Regenia albogularis. Proc. Zool. Soc. London 1861: 109-113 - get paper here
  • Haacke, W.D. 1995. Life History Notes - Varanus niloticus niloticus. African Herp News (22): 45-46 - get paper here
  • Haagner, G. V., Hall, R. L., & Branch, W. R. 1993. Varanidae: Varanus niloticus niloticus. J. Herp. Assoc. Africa, 42 Sept (38). - get paper here
  • Haagner, G.V.; Hall, R.; Branch, W.R. 1993. Life History Notes - Varanus niloticus niloticus. J. Herp. Assoc. Africa (42): 38-38 - get paper here
  • Hellmich, W. 1957. Herpetologische Ergebnisse einer Forschungsreise in Angola. Veröff. Zool. Staatssammlung München 5: 1-91 - get paper here
  • Hennessy, J. 2010. Parthenogenesis in an Ornate Nile Monitor, Varanus ornatus. Biawak 4 (1): 26-30 - get paper here
  • Hennig, A.S. 2006. Freilandbeobachtungen an Siedleragamen (Agama agama) und Nilwaranen (Varanus niloticus) in Senegal, Westafrika. Reptilia (Münster) 11 (61): 3-5 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Herrmann, Hans-Werner; Schmitz, Andreas; Herrmann, Patricia A. & Böhme, Wolfgang 2007. Amphibians and reptiles of the Tchabal Mbabo mountains, Adamaoua plateau, Cameroon. Bonner zoologische Beiträge 55 (1/2): 27-35 - get paper here
  • Hinsley, A. E., Willis, J., Dent, A., Oyanedel, R., Kubo, T., & Challender, D. 2023. Trading species to extinction: evidence of extinction linked to the wildlife trade. Cambridge Prisms: Extinction - get paper here
  • Hirth, H.F. & E.M.A. Latif 1979. Deep Body Temperatures of the Nile Monitor (Varanus niloticus) taken by Radiotelemetry Journal of Herpetology 13 (3): 367-368. - get paper here
  • Hoogmoed, M.S. 1980. Herpetologische waarnemingen in Ghana [part 5]. Lacerta 38 (7): 62-65 - get paper here
  • Iannucci, Alessio; Marie Altmanová, Claudio Ciofi, Malcolm Ferguson-Smith, Massimo Milan, Jorge Claudio Pereira, James Pether, Ivan Rehák, Michail Rovatsos, Roscoe Stanyon, Petr Velenský, Petr Ráb, Lukáš Kratochvíl & Martina Johnson Pokorná 2019. Conserved sex chromosomes and karyotype evolution in monitor lizards (Varanidae). Heredity 123, 215–227 - get paper here
  • ICZN 1959. Opinion 540. Protection under the plenary power of the specific names bengalensis Daudin, [1802], as published in the combination Tupinambis bengalensis, and salvator Laurenti 1768, as published in the combination Stellio salvator. Opin. Declar. intern. Com. zool. Nom. 20: 77-85
  • JACOBSEN, N.H.G. 2009. A contribution to the herpetofauna of the Passendro Area, Central African Republic. African Herp News (47): 2-20 - get paper here
  • Jacobsen, Niels H.G.; Errol W. Pietersen & Darren W. Pietersen 2010. A preliminary herpetological survey of the Vilanculos Coastal Wildlife Sanctuary on the San Sebastian Peninsula, Vilankulo, Mozambique. Herpetology Notes 3: 181-193
  • KEITH, M. & GINSBERG, A.E. 2010. Varanus niloticus (Linneaus, 1758) Feeding behaviour. African Herp News (51): 19-21 - get paper here
  • Kirchhof S, Wasonga V, Mazuch T, Spawls S, Malonza KP. 2023. An annotated checklist of the herpetofauna of the Sibiloi National Park in northern Kenya based on field surveys. Amphibian & Reptile Conservation 17(1/2) [General Section]: 1–18 (e324) - get paper here
  • Klitscher, Tom. 2014. Herpetologische Beobachtungen in Sambia, Simbabwe, Malawi und Tansania. Sauria 36 (1): 3-8 - get paper here
  • Lang, D.; Branch, W.R. 1990. Life History Note: Varanus niloticus niloticus: Diet. J. Herp. Assoc. Africa (37): 50-50 - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988] - get paper here
  • Largen, M.J.; Spawls, S. 2006. Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification. Tropical Zoology 19 (1): 21-109 - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Leaché, Adam D.; Mark-Oliver Rödel, Charles W. Linkem, Raul E. Diaz, Annika Hillers, and Matthew K. Fujita 2006. Biodiversity in a forest island: reptiles and amphibians of the Togo Hills, Kyabobo National Park, Ghana. Amphibian & Reptile Conservation 4 (1): 22-45 - get paper here
  • LeBreton, Matthew 1999. A working checklist of the herpetofauna of Cameroon. Netherlands Committee for IUCN, 160 pp.
  • Lenz, Sigrid 1995. Zur Biologie und Ökologie des Nilwarans, Varanus niloticus (Linnaeus 1766) in Gambia, Westafrica. Mertensiella 5: 1-256
  • Leptien, Rolf 2014. Auf der Suche nach dem Togo-Chamäleon. Draco 15 (58): 54-59 - get paper here
  • Linnaeus, C. [= Linné, C. von] 1758. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Holmiæ. 10th Edition: 824 pp. - get paper here
  • Lisle, H.F. de 1996. Natural History of Monitor Lizards. Krieger, Malabar (Florida)
  • Lobón-Rovira J, Vaz Pinto P, S. Becker F, Tolley KA, Measey J, Bennet B, Boon B, de Sá S, Conradie W 2022. An updated herpetofaunal species inventory of Iona National Park in southwestern Angola. Check List 18(2): 289-321 - get paper here
  • Loveridge, A. 1923. Notes on East African lizards collected 1920-1923 with the description of two new races of Agama lionotus Blgr. Proc. Zool. Soc. London 1923: 935-969 - get paper here
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Loveridge, A. 1938. On a collection of reptiles and amphibians from Liberia. Proc. New England Zool. Club 17: 49-74
  • Lyakurwa, J.V., Howell, K.M., Munishi, L.K., Treydte, A.C. 2019. Uzungwa Scarp Nature Forest Reserve; a unique hotspot for reptiles in Tanzania. Acta Herpetologica 14 (1): 3-14 - get paper here
  • Lyakurwa, John Valentine 2017. The Reptiles of the Uzungwa Scarp Forest Reserve (USFR): An Updated Checklist with Notes on Dagger-Tooth Vine Snake Xyelodontophis uluguruensis. Journal of East African Natural History 106(2):57-65. - get paper here
  • Malonza, Patrick K.; David M. Mulwa, Joash O. Nyamache, Georgina Jones 2017. Biogeography of the Shimba Hills ecosystem herpetofauna in Kenya. Zoological Research 38(5): 1-11 - get paper here
  • Malonza, Patrick K.; Victor D. Wasonga, Vincent Muchai , Damaris Rotich, Beryl A. Bwong; A.M. Bauer 2006. DIVERSITY AND BIOGEOGRAPHY OF HERPETOFAUNA OF THE TANA RIVER PRIMATE NATIONAL RESERVE, KENYA. Journal of East African Natural History 95(2): 95–109 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Mazzotti, F.J., J.H. Nestler, J.M. Cole, C. Closius, W.H. Kern, M.R. Rochford, E. Suarez, R. Brubaker, S.G. Platt, T. Rainwater & J.K. Ketterlin 2020. Diet of Nile Monitors (Varanus niloticus) Removed from Palm Beach and Broward Counties, Florida, USA Journal of Herpetology 54 (2): 189-195. - get paper here
  • Mármol, Gabriel Martínez del; Ander Izagirre-Egaña, Sonke Frahm & Marco Sassoè 2023. Herpetofauna of Mauritania: results of a field survey. Bol. Asoc. Herpetol. Esp. (2023) 34 Preprint-101 - get paper here
  • Meerman, J.C. 1979. Enkele herpetologische waarnemingen in Tanzania. Lacerta 37 (10-11): 154-163 - get paper here
  • Meier, Axel 2023. Uwemajo – ein Reptilienzoo zum Erhalt des Mangrovenwaldes auf Sansibar. Elaphe 2023 (4): 52-61
  • Meierkord, R. 2007. Amputation der Schwanzspitze von Varanus ornatus (DAUDIN 1803). Elaphe 15 (2): 38-40
  • Mertens, R. 1942. Die Familie der Warane (Varanidae), 1. Teil: Allgemeines. Abh. Senckenb. naturf. Ges., 462: 1-116.
  • Meshaka Jr., Walter E. 2011. A RUNAWAY TRAIN IN THE MAKING: THE EXOTIC AMPHIBIANS, REPTILES, TURTLES, AND CROCODILIANS OF FLORIDA. Herp. Cons. Biol. 6 (Monograph 1): 1-101 - get paper here
  • Meyer, F.A.A. 1795. Synopsis Reptilium, novam ipsorum sistens generum methodum, nec non Gottingensium huius ordinis animalium enumerationem. Vandenhoeck und Ruprecht, Göttingen, 32 pp [for current names see Rösler & Böhme 2021] - get paper here
  • Miguel, A. R., Kaiser, H., Vargas, N. D., & Borges-Martins, M. 2024. Diversity and natural history of reptiles from coastal environments in Gilé National Park, northern Mozambique. Herpetology Notes, 17, 173-199
  • Mitchell, A.J.L.; Steyn, W. 1967. Further distribution records of Reptiles in SW- Africa. Cimbebasia 21: 23-26 (Windhoek)
  • MOHAMEDIEN, DALIA & MAHMOUD AWAD. 2021. Pulmonary Guardians and Special Regulatory Devices in the Lung of Nile Monitor Lizard (Varanus niloticus) with Special Attention to the Communication Between Telocyte, Pericyte, and Immune Cells. Microscopy and Microanalysis (2021.): 1–7. - get paper here
  • MOHAMMED, ELSAYED S.I.; FATMA A. MADKOUR & RASHA RADEY. 2021. Histological features of the spleen and pancreas of the Egyptian Nile monitor lizard (Varanus niloticus). SVU-International Journal of Veterinary Sciences 4(4): 22–32. - get paper here
  • Monard, ALBERT 1937. Contribution à l'herpétologie d'Angola. Arq. Mus. Bocage, Lisbon 8:19-153.
  • Monasterio, Camila 2016. The herpetofauna of the Dindefelo Natural Community Reserve, Senegal. Herpetology Notes 9: 1-6 - get paper here
  • Musah, Yahaya; Benjamin Y. Ofori and Daniel K. Attuquayefio 2019. Herpetofauna community diversity and composition of a changing coastal wetland in Ghana. West African Journal of Applied Ecology 27 (1): 52 - 65
  • Nneji, Lotanna Micah; Adeniyi Charles Adeola, , Agboola Okeyoyin, Ojo Cornelius Oladipo, Yohanna Saidu, Dinatu Samuel, John Yinamga Usongo, Babatunde Emmanuel Adedeji, Olatunde Omotoso, Akindele OluwatosinAdeyi, ObihAlexUgwumba,andAdiahaAldaAlexUgwum 2019. Diversity and distribution of amphibians and reptiles in Gashaka Gumti National Park, Nigeria. Herpetology Notes 12: 543-559 - get paper here
  • Ollonen, J., Khannoon, E. R., Macrì, S., Vergilov, V., Kuurne, J., Saarikivi, J., ... & Di-Poï, N. 2024. Dynamic evolutionary interplay between ontogenetic skull patterning and whole-head integration. Nature Ecology & Evolution, 1-16 - get paper here
  • Padial, J. M. 2006. COMMENTED DISTRIBUTIONAL LIST OF THE REPTILES OF MAURITANIA (WEST AFRICA). Graellsia, 62(2): 159-178 - get paper here
  • Pauwels O. S.G., Carlino P., Chirio L., Daversa D.R., Lips J., Oslisly Richard, Testa O. 2019. Amphibians and reptiles found in caves in Gabon, western Equatorial Africa. Cave and Karst Science, 46 (1): 3-12 - get paper here
  • Pauwels, O. S. G. B. Le Garff, I. Ineich, P. Carlino, I. Melcore, L. Boundenga, C. Vigna, T. Stévart, K. Jeffery, C. Orbell, J.-B. Squarcini, J. P. Vande weghe and L. J. T. White 2016. Miscellanea Herpetologica Gabonica V & VI. Bull. Chicago Herp. Soc. 51: 177 - get paper here
  • Pauwels, O.S.G. & Vande Weghe, J.P. 2008. Les reptiles du Gabon. Smithsonian Institution, Washington: 272 pp. - get paper here
  • Pauwels, O.S.G., Albert, J.L.; Vande Weghe, G. & Gramentz, D. 2007. Neue Reptiliennachweise von Franceville, Südost-Gabun. Elaphe 15 (3): 63-66 - get paper here
  • Pauwels, O.S.G.; David, P. 2008. Miscellanea Herpetologica Gabonica I. Hamadryad 32 (1): 13 – 18 - get paper here
  • Pauwels, O.S.G.; Kamdem Toham, A. & Chimsunchart, C. 2002. Recherches sur l’herpétofaune du Massif du Chaillu, Gabon. Bull. Inst. Roy. Sci. Nat. Belgique (Biologie) 72: 47-57 - get paper here
  • Pauwels, O.S.G.; William R. Branch and Marius Burger 2004. REPTILES OF LOANGO NATIONAL PARK, OGOOUÉ-MARITIME PROVINCE, SOUTH-WESTERN GABON. Hamadryad 29 (1): 115 – 127 - get paper here
  • Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press, 599 pp - get paper here
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Portik DM, Jongsma GFM, Kouete MT, Scheinberg LA, Freiermuth B, Tapondjou WP, Blackburn DC. 2016. A survey of amphibians and reptiles in the foothills of Mount Kupe, Cameroon. Amphibian & Reptile Conservation 10(2) [Special Section]: 37–67 (e131)
  • Rese, R. 1988. Beitrag zur Lebenserwartung von Grosswaranen in Gefangenschaft. Sauria 10 (1): 25-26 - get paper here
  • Rieppel, O. & L Labhardt 1979. Mandibular mechanics in Varanus niloticus Herpetologica 35 (2): 158-163. - get paper here
  • Robertson, I. A. D., B. M. Chapman and N. F. Chapman 1963. Notes on some reptiles collected in the Rukwa Valley, S. W. Tanganyika. Ann. Mag. Nat. Hist. Ser. 13, 5(55): 421‑432 - get paper here
  • Rödel,M. O., Grabow,K., Hallermann,J. & Böckheler,C. 1997. Die Echsen des Comoé-Nationalparks, Elfenbeinküste. Salamandra 33 (4): 225-240 - get paper here
  • Rösler, Herbert & Wolfgang Böhme 2021. Das Leben von Friedrich Albrecht Anton Meyer (1768 – 1795) und die Herpetologie in seinen Schriften, unter besonderer Berücksichtigung der „Synopsis Reptilium“. Sekretär 21: 83-129
  • SÁNCHEZ-VIALAS, A., CALVO-REVUELTA, M. & DE LA RIVA, I. 2022. Synopsis of the terrestrial Reptiles of Equatorial Guinea. Zootaxa 5202 (1): 1-197 - get paper here
  • Schleicher, Alfred 2015. Reptilien Namibias. Namibia Scientific  Society, 276 pp.
  • Schleicher, Alfred 2020. Reptiles of Namibia. Kuiseb Publishers, Windhoek, Namibia, 271 pp.
  • Schlüter, Uwe 2010. Ernährung nord- und westafrikanischer Warane in der Natur und bei Terrarienhaltung. Reptilia (Münster) 15 (86): 36-45 - get paper here
  • Schmidt, Karl Patterson 1919. Contributions to the Herpetology of the Belgian Congo based on the Collection of the American Congo Expedition, 1909-1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bull. Amer. Mus. Nat. Hist. 39 (2): 385-624 - get paper here
  • Schmitz A., Euskirchen, O. & Böhme W. 2000. Zur Herpetofauna einer montanen Regenwaldregion in SW-Kamerun (Mt. Kupe und Bakossi-Gergland).III. Einige bemerkenswerte Vertreter der Familien Lacertidae, Scincidae, Varanidae, Elapidae und Viperidae. Herpetofauna 22 (124): 16-27 - get paper here
  • Segniagbeto, G.H.; Dekawole, J.K.; Ketoh, G.K.; Dendi, D.; Luiselli, L. 2022. Herpetofaunal Diversity in a Dahomey Gap Savannah of Togo (West Africa): Effects of Seasons on the Populations of Amphibians and Reptiles. Diversity 14: 964
  • Segniagbeto, Gabriel Hoinsoude; Jean-François Trape, Komlan M. Afiademanyo, Mark-Oliver Rödel, Annemarie Ohler, Alain Dubois, Patrick David, Danny Meirte, Isabelle Adolé Glitho, Fabio Petrozzi, and Luca Luiselli 2015. Checklist of the lizards of Togo (West Africa), with comments on systematics, distribution, ecology, and conservation. Zoosystema 37 (2): 381-402 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sow, Andack Saad; Duarte Vasconcelos Gonçalves, Fábio Vieira Sousa, Fernando Martínez- Freiría, Frederico Santarém, Guillermo Velo-Antón, Hamidou Dieng, João Carlos Campos, Saer Khayar Diagne, Zbyszek Boratyński, José Carlos Brito 2017. Atlas of the distribution of amphibians and reptiles in the Diawling National Park, Mauritania. Basic and Applied Herpetology - get paper here
  • Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Sternfeld, R. 1917. Reptilia und Amphibia. In: Schubotz, H. (Hrsg.): Wissenschaftliche Ergebnisse der Zweiten Deutschen Zentral-Afrika-Expedition, 1910-1911 unter Führung Adolph Friedrichs, Herzog zu Mecklenburg. Leipzig: Klinkhardt & Biermann, [Band] 1, Zoologie, Lieferung 11; S. 407-510. - get paper here
  • Switak, Karl H. 1997. Varanids I have known: The Nile Monitor, Varanus niloticus. Monitor: Journal of the Victorian Herpetological Society 8 (3): 177-179
  • Thieme, U. 2004. Herpetologische Beobachtungen und anderes aus Gambia. Reptilia (Münster) 9 (49): 43-48 [erratum in 9 (6): 17] - get paper here
  • Tolley KA, Telford NS, Makhubo BG, Power RJ, Alexander GJ 2023. Filling the gap: Noteworthy herpetological discoveries in North West Province, South Africa. Zoosystematics and Evolution 99(1): 101-116 - get paper here
  • Trape, J.F.; Trape, S. & Chirio, L. 2012. Lézards, crocodiles et tortues d'Afrique occidentale et du Sahara. IRD Orstom, 503 pp. - get paper here
  • Ullenbruch, K.; Grell, O.; Böhme, W. 2010. Reptiles from southern Benin, West Africa, with the description of a new Hemidactylus (Gekkonidae), and a country-wide checklist. Bonn Zool. Bull. 57 (1): 31-54 - get paper here
  • Venter, Jan A.; Werner Conradie 2015. A checklist of the reptiles and amphibians found in protected areas along the South African Wild Coast, with notes on conservation implications. Koedoe 57 (1): 1-25. doi: 10.4102/koedoe.v57i1.1247 - get paper here
  • Werner,F. 1919. Wissenschaftliche Ergebnisse der mit Unterstützung der Kaiserlichen Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Aegyptischen Sudan (Kordofan) 1914. IV. Bearbeitung de Denkschr. Akad. Wiss Wien, Math.-Naturw. Klasse 96: 437-509 - get paper here
  • Witte, G. F. de 1933. Reptiles récoltés au Conge Belge par le Dr. H. Schouteden et par M. G.-F. Witte. Ann. Mus. Conge belge Zool. Ser. 1 Tome III: 53-100.
  • Wood, Jared P.; Todd S. Campbell and Robert B. Page 2016. Characterization of 17 novel microsatellite loci in the Nile monitor (Varanus niloticus) via 454 pyrosequencing. Amphibia-Reptilia 37 (1): 121 - 125 - get paper here
  • Wright, Kevin 1993. Nile monitors. Reptile & Amphibian magazine 1993 (Sept/Oct.): 36-44
  • Yeboah, S. 1994. Aspects of the biology of two sympatric species of monitor lizards Varanus niloticus and Varanus exanthematicus (Reptilia, Sauria) in Ghana. African Journal of Ecology 32 (4): 331-333 - get paper here
 
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