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Acanthodactylus scutellatus (AUDOUIN, 1827)

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesAcanthodactylus scutellatus audouini BOULENGER 1918
Acanthodactylus scutellatus scutellatus (AUDOUIN 1827) 
Common NamesE: Nidua Fringe-fingered Lizard, Nidua lizard, Nidua spiny-footed lizard 
SynonymLacerta scutellata AUDOUIN 1827: 117
Lacerta scutellata — MILNE-EDWARDS 1829: 85
Acanthodactylus scutellatus — DUMÉRIL & BIBRON 1839: 272
Acanthodactylus inornatus GRAY 1845: 38 (fide BOULENGER 1881: 744)
Acanthodactilus scutellatus — ZULUETA 1909: 355
Acanthodactylus scutellatus — BOULENGER 1918
Acanthodactylus scutellatus var. audouini BOULENGER 1918
Acanthodactylus scutellatus audouini — BONS & GIROT 1964
Acanthodactylus inornata — BONS & GIROT 1964
Acanthodactylus scutellatus — SALVADOR 1982
Acanthodactylus scutellatus — SCHLEICH, KÄSTLE & KABISCH 1996: 397
Acanthodactylus scutellatus — HARRIS & ARNOLD 2000
Acanthoiadylus scutellatus — HABEEB 2018 (in error)
Acanthodactylus scutellatus — BAR et al. 2021

Acanthodactylus scutellatus audouini BOULENGER 1918
Acanthodactylus scutellatus audouini — CROCHET et al. 2003
Acanthodactylus scutellatus audouini — SCHLÜTER 2010
Acanthodactylus scutellatus audouini — TRAPE, CHIRIO & TRAPE 2012 
DistributionMorocco ?, SE Algeria, S Tunisia, Libya, Egypt, Israel, Jordan, NE Mali, N Niger, N Chad, N Sudan (Jumhūriyyat), Iraq, N Saudi Arabia, Kuwait

audouini: Distributed in the west of the species range, Mauritania, Algeria, Tunisia, Egypt, Mali, to NE Sudan (Bons & Girot, 1964). Type locality: “Egypte, Nubie, Tripoli, sud de la Tunisie”; Type locality: Wed El Kreil, Tunisia.

Type locality: “Egypte”  
TypesHolotype: lost (fide CROCHET et al. 2003).
Lectotype: BMNH 1920.1.20.3006 designated by CROCHET et al. 2003. Paralectotypes: BMNH (Wadi Halfa, Sudan), BMNH 1913.12.30.6-10 (Homs, Tripoli), BMNH (Duirat, southern Tunisia), BMNH 1920.1.20.3006 (Wed El Kreil, Tunisia); see also SALVADOR 1982 [audouini] 
DiagnosisDiagnosis: Elongated pointed snout. Generally three supraoculars. One row of granules between the supraoculars and the superciliaries. Five supralabials anterior to the subocular. Small, keeled temporals. Tympanic scale absent. First three pairs of submaxillaries in contact. Dorsals small, flat, granular, and weakly keeled in relatively high number. 14 oblique longitudinal rows of ventrals. Fingers with four series of scales. Intense pectination on fourth toe. Reticulate color pattern. (Salvador 1982: 114). 
CommentSimilar species: Has been confused with A. longipes and A. dumerili.

Distribution: Erroneously reported from Morocco. See Bar et al. 2021 for a map.

Synonymy: SALVADOR (1982) synonymized A. inornata with A. scutellatus.

Date of publication: it is AUDOUIN 1827 (not 1809 or 1829 as sometimes stated; see remark in CROCHET et al. 2003: 141).

Subspecies: Acanthodactylus scutellatus hardyi HAAS, 1957 is now considered as a valid species. 
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