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Advenus montisilvestris (MYERS, 1973)

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Higher TaxaDiploglossidae, Diploglossa, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Mountain Forest galliwasp
G: Bergwald-Gallwespenschleiche 
SynonymDiploglossus montisilvestris MYERS 1973
Diploglossus montisilvestris — VILLA & OCCHIPINTI 1988
Diploglossus montisilvestris — KÖHLER 2000: 41
Advenus montisilvestris — SCHOOLS & HEDGES 2021: 217 
DistributionE Panama

Type locality: Panama, Darien, southeastern slope of Cerro Pirre, elevation 1440 m; 7.9242, - 77.7000 fide Schools & Hedges 2021.  
Reproductionoviparous 
TypesHolotype: KU 1113660 (field no. CWM 4995),an adult female caught by Charles W. Myers and Tomas Quintero on May 25, 1965. 
DiagnosisDiagnosis (genus). Species of Advenus gen. nov. have (1) claw sheath, present, (2) contact between the nasal and rostral scales, absent, (3) scales in contact with the nasal scale, four, (4) postnasal scales, one, (5) position of the nostril in the nasal scale, central, (6) keels on dorsal body scales, absent, (7) digits per limb, five, (8) longest toe lamellae, 16–17, (9) dorsal scale rows, 96, (10) relative head width, 12.0, (11) relative rostral height, 50.1, (12) relative frontonasal length, 2.46, (13) relative interparietal distance, 0.632, (14) relative axilla-groin distance, 60.0. From Caribicus gen. nov., we distinguish Advenus gen. nov. by the claw sheath (present versus its absence in Caribicus gen. nov.), keels on the dorsal scales (absent versus their presence in Caribicus gen. nov.), relative head width (12.0 versus 13.6–17.2), relative frontonasal length (2.46 versus 2.98–3.32), and the relative axilla-groin distance (60.0 versus 67.1–69.1). From Celestus, we distinguish Advenus gen. nov. by the claw sheath (present versus its absence in Celestus) and the relative axilla-groin distance (60.0 versus 60.9–66.3). From Comptus gen. nov., we distinguish Advenus gen. nov. by the claw sheath (present versus its absence in Comptus gen. nov.), keels on dorsal scales (absent versus their presence in Comptus gen. nov.), relative rostral height (50.1 versus 53.2–65.5), relative frontonasal length (2.46 versus 2.95–3.65), and the relative interparietal distance (0.632 versus 0–0.435). From Panolopus, we distinguish Advenus gen. nov. by the claw sheath (present versus its absence in Panolopus) and the relative axilla-groin distance (60.0 versus 49.7–59.6). From Sauresia, we distinguish Advenus gen. nov. by the digits per limb (five versus four), the longest toe lamellae (16–17 versus 8–12), the dorsal scale rows (96 versus 101–127), the relative interparietal distance (0.632 versus 0–0.431), and the relative axilla-groin distance (60.0 versus 63.9–69.9). From Wetmorena, we distinguish Advenus gen. nov. by the digits per limb (five versus four), the number of longest toe lamellae (16–17 versus 8–12), and the dorsal scale rows (96 versus 98–117). (Schools & Hedges 2021). 
CommentType species: Diploglossus montisilvestris MYERS 1973 is the type species of the genus Advenus SCHOOLS & HEDGES 2021. The genus is monotypic.

Abundance: Known from only one specimen collected on the southeastern slope of Cerro Pirre (1440 m), Serranía de Pirre, Province of Darién, Republic of Panama. 
EtymologyThe specific epithet means "of mountain forest," and is derived from the Latin prefix monti- (from mons, mountain) plus the adjective silvestris (belonging to forest).

The genus name Advenus is a masculine noun derived from the Latin advena (“stranger”), referring to the distribution of this species in Middle America when all of its close relatives are on Caribbean islands. 
References
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • Myers, C.W. 1973. Anguid lizards of the genus Diploglossus in Panamá, with the description of a new species. American Museum Novitates (2523): 1-20 - get paper here
  • SCHOOLS, MOLLY & S. BLAIR HEDGES 2021. Phylogenetics, classification, and biogeography of the Neotropical forest lizards (Squamata, Diploglossidae). Zootaxa 4974 (2): 201–257 - get paper here
  • Villa J D; Occhipinti N A 1988. Diploglossus montisilvestris Myers. Catalogue of American Amphibians and Reptiles ( 444: 1 - get paper here
 
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