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Agama atra DAUDIN, 1802

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Southern African Rock Agama 
SynonymAgama atra DAUDIN 1802: 349
Agama ater — GRAY 1831
Saura spinalis (partim) WAGLER 1833 (nom. subst. pro Agama atra)
Agama atra — DUMÉRIL & BIBRON 1837: 493
Phrynopsis atra — FITZINGER 1843f
Agama atra — BOULENGER 1885: 352
Agama atra — BOETTGER 1886: 12
Agama micropholis MATSCHIE 1891
Agama microterolepis BOULENGER 1896: 22
Agama holubi BOCAGE 1896 (syn. fide FITZSIMONS 1943)
Agama atra — ROUX 1907: 415
Agama holubi — GOUGH 1909
Agama atra — BOULENGER & POWER 1921: 273
Agama atra var. rudis BOULENGER & POWER 1921: 282
Agama atra atra — AUERBACH 1987: 96
Agama atra — MACEY et al. 2000
Agama atra — BATES et al. 2014: 306
Agama atra — SCHLEICHER 2020 
DistributionRepublic of South Africa (Eastern Cape), Swaziland, S Namibia, S Botswana

Type locality: unknown. Designated as “South Africa” by FITZSIMONS 1943.  
Reproductionoviparous 
TypesSyntypes: MNHN-RA 2604-2605, MNHN-RA 2604A and MNHN-RA 2605A (syntypes reported from BMNH are in error, fide I. Ineich, pers. comm.)
Holotype: BMNH 1946.8.27.94-96 (95.10.19.57) [microterolepis]
Lectotype: ZMB 42887 (designated by DENZER et al. 1997), ad. Ex., "Mpome", leg. A. & M. Krothe. Paralectotype: ZMB 42888, juv. Ex., same as lectotype; Terra typica: Mpome, Transvaal [micropholis] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (85 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy partly after WERMUTH 1967 and P. Wagner (pers. comm., 24 Oct 2011).

Subspecies: MATTHEE & FLEMMING 2002 and WAGNER et al. (2009) elevated A. a. knobeli to full species status. 
EtymologyNamed after its color, Latin “ater, atra, atrum” = dark or black. 
References
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
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  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001]
  • Boettger, O. 1886. Beitrag zur Herpetologie und Malakozoologie Südwest-Afrikas. 1. Zur Kenntniss der Fauna von Angra Pequenia. Abh. senckenb. naturf. Ges. (Frankfurt) 1886: 3-29 - get paper here
  • Boulenger, G. A. 1896. Descriptions of two new lizards from the Transvaal. Ann. Mag. nat. Hist. (6) 17: 21—22 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
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  • Boulenger,G.A. & POWER, J.H. 1921. A revision of the South African agamas allied to Agama hispida and A. atra. Transact. roy. Soc. S. Africa, CapeTown, 9: 229-287; https://doi.org/10.1080/00359192109520217 - get paper here
  • Branch, W. R. 1988. Life History Note: Agama atra atra: Reproduction. J. Herp. Assoc. Africa (35): 39-39 - get paper here
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  • Conradie, Werner; Brain Reeves, Sandile Mdoko, Lwandiso Pamla, Oyama Gxabhu 2020. Herpetological survey of the Ongeluksnek (Malekgalonyane) Nature Reserve on the foothills of the Drakensberg, Eastern Cape Province, South Africa. Herpetology Notes 13: 717-730 - get paper here
  • Conradie, Werner; Brian Reeves , Neil Brown, & Jan A. Venter 2016. Herpetofauna of the Oviston, Commando Drift and Tsolwana nature reserves in the arid interior of the Eastern Cape Province, South Africa. Indago 32: 81-98 - get paper here
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