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Agama aculeata MERREM, 1820

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards) 
SubspeciesAgama aculeata distanti (BOULENGER 1902)
Agama aculeata aculeata MERREM 1820 
Common NamesGround Agama
aculeata: Western Ground Agama
distanti: Northern Ground Agama 
SynonymAgama aculeata MERREM 1820: 53
Agama aculeata DUMÉRIL & BIBRON 1837: 499 (non MERREM ?)
Trapelus (Psammorrhoa) bibronii FITZINGER 1843 (nom. subst.)
Saura spinalis WAGLER 1866 (nom. subst., partim)
Agama infralineata PETERS 1877
Agama aculeata — BOULENGER 1885: 351
Agama pulchella BOCAGE 1896
Agama hispida aculeata — LOVERIDGE 1936: 52
Agama hispida aculeata — FITZSIMONS & BRAIN 1958
Agama aculeata aculeata — AUERBACH 1987: 98
Agama aculeata — PIANKA & VITT 2003: 123
Agama aculeata aculeata — BATES et al. 2014: 303

Agama aculeata distanti (BOULENGER 1902)
Agama distanti BOULENGER 1902: 339
Agama hispida var. distanti — LOVERIDGE 1923: 942
Agama aculeata distanti — BOYCOTT 1992
Agama aculeata distanti — BATES et al. 2014: 30 
DistributionNamibia, Botswana, Zimbabwe, Republic of South Africa, Mozambique, S Angola, Zambia, Swaziland

Type locality: Otjimbingue [Namibia] [Agama infralineata PETERS 1877]

distanti: Republic of South Africa (Limpopo, Mpumalanga, Gauteng, NW Province, Free State, Kwazulu-Natal), Swaziland  
Reproductionoviparous 
TypesHolotype: ZMB 750, "in promontorio bonae spei", leg. v. Borcke (actually a Paralectotypus fide TILLACK, pers. comm. 22 Jan 2014, Wagner et al. 2012)
Lectotype: Fig. 7 of Seba (1735, II, tomus 8)
Holotype: MNHN [Agama aculeata DUMÉRIL & BIBRON 1837]
Syntypes: ZMB 4217, 4218, 4219 [Agama infralineata PETERS 1877] 
Diagnosis 
CommentAgama aculeata armata is treated here as a full species.

Synonymy: WERMUTH 1965, WAGNER et al. 2012.

Distribution: not in East Africa; P. Wagner suggested to restrict this name to South African populations (pers. comm. to S. Spawls, 28 June 2018). 
Etymology 
References
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Barts, M. 2003. Die Agamen des südlichen Afrikas. Draco 4 (14): 70-79 - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bauer, Aaron M.; Branch, William R. & Haacke, Wulf D. 1993. The herpetofauna of the Kamanjab area and adjacent Damaraland, Namibia. Madoqua (Windhoek) 18 (2): 117-145.
  • Billawer, W.H.; Heidemann, N.J.L. 1996. A comparative analysis of diurnal behavioural activities in males of Agama aculeata aculeata and Agama planiceps planiceps in Windhoek, Namibia. J. Herpet. Assoc. Africa 45 (2): 68-73 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A. 1902. A new name for the Common Agama of the Transvaal. Ann. Mag. Nat. Hist. (7) 9: 339-339 - get paper here
  • Boycott, R.C. 1992. An Annotated Checklist of the Amphibians and Reptiles of Swaziland. The Conservation Trust of Swaziland - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Cimatti, E. 2007. Namibia - Introduction to a vast territory. Reptilia (GB) (50): 58-67 - get paper here
  • Conradie W, Bills R, and Branch WR. 2016. The herpetofauna of the Cubango, Cuito, and lower Cuando river catchments of south-eastern Angola. Amphibian & Reptile Conservation 10(2) [Special Section]: 6–36 - get paper here
  • CONRADIE, W., DOUCETTE-RIISE, S., VANHOOYDONCK, B., ENGELBRECHT, H., MEASEY, G.J., & TOLLEY, K. 2011. Herpetological survey of Rooipoort Nature Reserve, Northern Cape, South Africa. African Herp News (53): 35-41
  • Conradie, Werner; Brian Reeves , Neil Brown, & Jan A. Venter 2016. Herpetofauna of the Oviston, Commando Drift and Tsolwana nature reserves in the arid interior of the Eastern Cape Province, South Africa Indago 32: 81-98 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Fitzsimons, V.F.M.; Brain, C.K. 1958. A Short account of the Reptiles of the Kalahari Gemsbok National Park. Koedoe, 1(1): 99-104 - get paper here
  • Haacke, W. D. 1984. The herpetology of the southern Kalahari domain. In: Proc. Symp. Kalahari Ecosystems. Koedoe, Suppl., 1984: 171-186 - get paper here
  • Heideman, N.J.L. 1993. Social Organization and Behaviour of Agama aculeata aculeata and Agama planiceps planiceps during the Breeding Season. J. Herp. Assoc. Africa (42): 29-31 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • LEACHÉ, Adam D.; Rebecca A. CHONG, Theodore J. PAPENFUSS, Philipp WAGNER, Wolfgang BÖHME, Andreas SCHMITZ, Mark-Oliver RÖDEL, Matthew LEBRETON, Ivan INEICH, Laurent CHIRIO, Aaron BAUER, Edem A. ENIANG, & Sherif BAHA EL DIN 2009. Phylogeny of the genus Agama based on mitochondrial DNA sequence data. Bonner zoologische Beiträge 56 (4): 273–278 - get paper here
  • Loveridge, A. 1923. Notes on East African lizards collected 1920-1923 with the description of two new races of Agama lionotus Blgr. Proc. Zool. Soc. London 1923: 935-969 - get paper here
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • McLachlan G R 1981. Taxonomy of Agama hispida (Sauria: Agamidae) in southern Africa. Cimbebasia Ser. A 5(6): 219-227
  • Merrem, B. 1820. Versuch eines Systems der Amphibien I (Tentamen Systematis Amphibiorum). J. C. Kriegeri, Marburg, 191 pp.
  • Peters, Wilhem Carl Hartwig 1877. Übersicht der Amphibien aus Chinchoxo (Westafrika), welche von der Africanischen Gesellschaft dem Berliner zoologischen Museum übergeben sind. Monatsber. königl. Akad. Wiss. Berlin. 1877 (October): 611-620 - get paper here
  • Pianka, E.R. & Vitt, L.J. 2003. Lizards - Windows to the Evolution of Diversity. University of California Press, Berkeley, 347 pp. [review in Copeia 2004: 955] - get paper here
  • Schleicher, Alfred 2015. Reptilien Namibias. Namibia Scientific  Society, 276 pp.
  • Wagner, Philipp; Aaron M. Bauer, Thomas M. Wilms, Mirko Barts, Wolfgang Böhme 2012. Miscellanea Accrodontia: Notes on Nomenclature, Taxonomy and Distribution. Russ. J. Herpetol. 19 (2): 177-189 - get paper here
 
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