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Agama atra DAUDIN, 1802

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Higher TaxaAgamidae (Agaminae), Sauria (lizards) 
Subspecies 
Common NamesSouthern Rock Agama 
SynonymAgama atra DAUDIN 1802: 349
Agama ater — GRAY 1831
Saura spinalis (partim) WAGLER 1833 (nom. subst. pro Agama atra)
Agama atra — DUMÉRIL & BIBRON 1837: 493
Phrynopsis atra — FITZINGER 1843
Agama atra — BOULENGER 1885: 352
Agama micropholis MATSCHIE 1891
Agama microterolepis BOULENGER 1896: 22
Agama atra rudis BOULENGER & POWER 1921
Agama atra atra — AUERBACH 1987: 96
Agama atra — MACEY et al. 2000 
DistributionRepublic of South Africa, Swaziland, S Namibia, S Botswana

Type locality: unknown. Designated as “South Africa” by FITZSIMONS 1943. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesType: BMNH 1946.8.27.94-96 (and possibly additional specimens).
Lectotype: ZMB 42887 (designated by DENZER et al. 1997), ad. Ex., "Mpome", leg. A. & M. Krothe. Paralectotype: ZMB 42888, juv. Ex., same as lectotype; Terra typica: Mpome, Transvaal [micropholis] 
CommentSynonymy partly after WERMUTH 1967 and P. Wagner (pers. comm., 24 Oct 2011).

Subspecies: MATTHEE & FLEMMING 2002 and WAGNER et al. (2009) elevated A. a. knobeli to full species status. 
EtymologyNamed after its color, Latin “ater, atra, atrum” = dark or black. 
References
  • Auerbach,R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Barts, M. 2003. Die Agamen des südlichen Afrikas. Draco 4 (14): 70-79 - get paper here
  • Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001]
  • Boulenger, G. A. 1896. Descriptions of two new lizards from the Transvaal. Ann. Mag. nat. Hist. (6) 17: 21—22 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Boulenger,G.A. & POWER, J.H. 1921. A revision of the South African agamas allied to Agama hispida and A. atra. Transact. roy. Soc. S. Africa, CapeTown, 9: 229-287
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Bruton & HAACKE 1980. Studies on the Ecology of Maputoland. 254 pp.
  • CONRADIE, W., DOUCETTE-RIISE, S., VANHOOYDONCK, B., ENGELBRECHT,<br />H., MEASEY, G.J., & TOLLEY, K. 2011. Herpetological survey of Rooipoort Nature Reserve, Northern Cape, South Africa. Afr. Herp News (53): 35-41
  • Cooper Jr., W.E. 2005. Duration of movement as a lizard foraging movement variable. Herpetologica 61 (4): 363-372 - get paper here
  • Daudin, F. M. 1802. Histoire Naturelle, génerale et particulièredes reptiles, ouvrage faisant suite, a l'histoiure naturelle, générale et particulière composée par LECLERC DE BUFFON, et redigée par C. S. SONNINI, vol. 3. F. Dufart, Paris.
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • FitzSimons, V. F. 1935. Notes on a collection of reptiles and amphibians made in the Southern Kalahari, Bushmanland, and Great and Little Namaqualand. Ann. Transvaal Mus., Pretoria 17: 519-550 - get paper here
  • Grubenmann, M. 2013. Einige fotografische Beobachtungen an Agamen in Kenia, Tansania, Malawi, Südafrika und Namibia. Iguana 26 (1): 23-33 [spelled as Grubermann on paper]
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • LEACHÉ, Adam D.; Rebecca A. CHONG, Theodore J. PAPENFUSS, Philipp WAGNER, Wolfgang BÖHME, Andreas S 2009. Phylogeny of the genus Agama based on mitochondrial DNA sequence data. Bonner zoologische Beiträge 56 (4): 273–278 - get paper here
  • Loveridge,A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Macey, J. R., J. A. Schulte II, A. Larson, N. B. Ananjeva, Y. Wang, R. Pethiyagoda, N. Rastegar-Pouy 2000. Evaluating trans-Tethys migration: an example using acrodont lizard phylogenetics. Systematic Biology 49 (2): 233-256 - get paper here
  • Matschie,G.F.P. 1891. Ueber eine kleine Sammlung von Reptilien und Amphibien aus Süd-Afrika. Zool. Jahrb., Abtl. Systematik (Jena) 5 [1890]: 605-611 - get paper here
  • Matthee, C.A. & Flemming, A.F., 2002. Population fragmentation in the southern rock agama, Agama atra: more evidence for vicariance in Southern Africa. Molecular Ecology 11: 465-471
  • Mertens,R. 1955. Die Amphibien und Reptilien Südwestafrikas. Aus den Ergebnissen einer im Jahre 1952 ausgeführten Reise. Abh. senckenb. naturf. Ges. (Frankfurt) 490: 1-172
  • Schönecker, P. 2014. Südafrika. Auf Reptiliensuche entlang der Garden Route. Reptilia (Münster) 19 (105): 80-93
  • Tolley, Krystal A.; Jane Sakwa Makokha, Darren T. Houniet, Belinda L. Swart and Conrad A. Matthee 2009. The potential for predicted climate shifts to impact genetic landscapes of lizards in the South African Cape Floristic Region. Molecular Phylogenetics and Evolution 51 (1): 120-130 - get paper here
  • van Wyk, J.H. & L. Ruddock2000 2001. The male reproductive cycle of the lizard Agama atra (Sauria: Agamidae) in central South Africa. Amphibia-Reptilia 22 (1): 119-123 - get paper here
  • Wagler,J. G. 1833. Isis von Oken 26: 896, 897 - get paper here
  • Wagner, P.; Ineich, I.; Leaché, A.D.; Wilms, T.M.; Trape, S.; Böhme, W. & Schmitz, A. 2009. Studies on African Agama VI. Taxonomic status of the West African Agama (Sauria: Agamidae) with prominent tail crests: Agama boulengeri Lataste 1886, Agama insularis Chabanaud, 1918 and Agama cristata Mocquard, 1905. Bonner zoologische Beiträge 56 (4): 239–253 - get paper here
 
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