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Agama atra DAUDIN, 1802

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Higher TaxaAgamidae (Agaminae), Sauria (lizards) 
SubspeciesAgama atra knobeli BOULENGER & POWER 1921
Agama atra atra DAUDIN 1802 
Common NamesSouthern Rock Agama 
SynonymAgama atra DAUDIN 1802: 349
Agama ater — GRAY 1831
Saura spinalis (partim) WAGLER 1833 (nom. subst. pro Agama atra)
Agama atra — DUMÉRIL & BIBRON 1837: 493
Phrynopsis atra — FITZINGER 1843
Agama atra — BOULENGER 1885: 352
Agama micropholis MATSCHIE 1891
Agama microterolepis BOULENGER 1896: 22
Agama atra rudis BOULENGER & POWER 1921
Agama atra — MACEY et al. 2000

Agama atra atra DAUDIN 1802
Agama atra atra — AUERBACH 1987: 96

Agama atra knobeli BOULENGER & POWER 1921
Agama anchietae var. knobeli BOULENGER & POWER 1921
Agama anchietae knobeli — FITZSIMONS 1935: 532
Agama atra knobeli — MERTENS 1955: 57
Agama atra knobeli — WERMUTH 1967: 8
Agama knobeli — WAGNER et al. 2009 
DistributionRepublic of South Africa, Swaziland, S Namibia, S Botswana

knobeli: Namibia (Mountains between Lüderitz Bay and Keetmanshoop). Type locality: SW Africa

Type locality: unknown. Designated as “South Africa” by FITZSIMONS 1943. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesType: BMNH 1946.8.27.94-96 (and possibly additional specimens).
Lectotype: ZMB 42887 (designated by DENZER et al. 1997), ad. Ex., "Mpome", leg. A. & M. Krothe. Paralectotype: ZMB 42888, juv. Ex., same as lectotype; Terra typica: Mpome, Transvaal [micropholis] 
CommentSynonymy partly after WERMUTH 1967 and P. Wagner (pers. comm., 24 Oct 2011).

Subspecies: WAGNER et al. (2009) elevated A. a. knobeli to full species status. However, their diagnosis is not sufficiently quantitative to make a clear distinction: “Agama atra is lacking the prominent large tail crest and has more spinose scales as A. knobeli, but further research on the relationships of these two species is in need.However, A. knobeli differs from A. cristata in (a) having larger body scales, (b) dorsal scales in same size as ventral scales and (c) in having a pale vertebral stripe in nuptial coloured adult males, and (d) a geographic distribution restricted to Namibia.” 
EtymologyEtymology: Named after its color, Latin “ater, atra, atrum” = dark or black. 
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  • Mertens,R. 1955. Die Amphibien und Reptilien Südwestafrikas. Aus den Ergebnissen einer im Jahre 1952 ausgeführten Reise. Abh. senckenb. naturf. Ges. (Frankfurt) 490: 1-172
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  • Wagner, P.; Ineich, I.; Leaché, A.D.; Wilms, T.M.; Trape, S.; Böhme, W. & Schmitz, A. 2009. Studies on African Agama VI. Taxonomic status of the West African Agama (Sauria: Agamidae) with prominent tail crests: Agama boulengeri Lataste 1886, Agama insularis Chabanaud, 1918 and Agama cristata Mocquard, 1905. Bonner zoologische Beiträge 56 (4): 239–253 - get paper here
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