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Alopoglossus viridiceps TORRES-CARVAJAL & LOBOS, 2014

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Higher TaxaAlopoglossidae, Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common NamesE: Green-headed shade lizards
S: Lagartijas de sombra de cabeza verde 
SynonymAlopoglossus viridiceps TORRES-CARVAJAL & LOBOS 2014
Alopoglossus viridiceps — HERNÁNDEZ-MORALES et al. 2020 
DistributionEcuador (Pichincha)

Type locality: Nanegal, Santa Lucia Cloud Forest Reserve, 0.113528°N; -78.6135°W (Decimal Degrees, WGS84), 1742 m, Provincia Pichincha, Ecuador  
Reproductionoviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) 
TypesHolotype: QCAZ 10670 (Figs 1,2), an adult male, collected on 27 June 2010 by V. Aguirre-Peñafiel and J. Zanka. Paratypes (11). ECUADOR: Provincia Pichincha: QCAZ 9738, Mindo, Haci- enda San Vicente, -0.050720°N, -78.772350°W (DD), 1246 m, collected on 7 Au- gust 2009 by S. Poe, E. Schaad, I. Latella, N. Blea, T. Kennedy and F. Ayala-Varela; QCAZ 10821, 10826, Nanegal, Santa Lucía Cloud Forest Reserve, 0.117780°N, -78.607555°W (DD), 1580 m, collected on 9 March 2010 by B. Tolhurst, P. Mafla- Endara, S. Ryan and X. Cueva; QCAZ 11854–55, Nanegal, Santa Lucía Cloud For- est Reserve, trail to waterfalls, 0.109450°N, -78.609380°W (DD), 1645 m, collect- ed on 12 September 2013 by D. Ortiz and O. Torres-Carvajal; QCAZ 11927–29, Nanegal, Santa Lucía Cloud Forest Reserve, 0.113330°N, -78. 613280°W (DD), 1736 m, collected on 6 November 2013 by F. Ayala-Varela, E. Carrillo, V. Macias and T. Ostos; QCAZ 10666, 10753, same collection data as holotype, but collected on 14 July 2010 by V. Aguirre-Peñafiel and 26 July 2010 by S. Maddock and V. Aguirre-Peñafiel, respectively. QCAZ 10671, Nanegal, Santa Lucía Cloud Forest Re- serve, 0.119280°N, -78.596470°W (DD), 1911 m, collected on 29 June 2010 by M.A. Chinchero. 
DiagnosisDiagnosis: Alopoglossus viridiceps can be distinguished from all other known con- geners except A. festae by having a double longitudinal row of widened gular scales and lanceolate dorsal scales in transverse rows. From A. festae (character states in paren- theses, taken from Köhler et al. 2012), the new species differs in having 29–32 dorsal scales in a transverse row at midbody (16–24, mean = 19.14 ± 2.25), four ventral scales in a transverse row at midbody (six), and a distinct longitudinal light stripe from mouth commissure to shoulder (Fig. 3). Scale counts and measurements of A. festae and A. viridiceps are presented in Table 2 in TORRES-CARVAJAL & LOBOS 2014. 
Comment 
References
  • Arteaga A, Pyron RA, Peñafiel N, Romero-Barreto P, Culebras J, Bustamante L, et al. 2016. Comparative Phylogeography Reveals Cryptic Diversity and Repeated Patterns of Cladogenesis for Amphibians and Reptiles in Northwestern Ecuador. PLoS One 11 (4): e0151746.doi:10.1371/journal.pone.0151746 - get paper here
  • Arteaga, A.; Bustamante, L.; Vieira, J. 2024. Reptiles of Ecuador. Khamai Foundation & Tropical Herping, 1073 pp. - get paper here
  • Hernández-Morales, C., Sturaro, M.J., Nunes, P.M.S., Lotzkat, S. and Peloso, P.L. 2020. A species‐level total evidence phylogeny of the microteiid lizard family Alopoglossidae (Squamata: Gymnophthalmoidea). Cladistics - get paper here
  • Ribeiro-Junior, M. A., Choueri, E., Lobos, S., Venegas, P., Torres-Carvajal, O. and Werneck, F. 2020. Eight in one: morphological and molecular analyses reveal cryptic diversity in Amazonian alopoglossid lizards (Squamata: Gymnophthalmoidea). Zoological Journal of the Linnean Society 190 (1): 227–270 - get paper here
  • Torres-Carvajal O, Lobos SE 2014. A new species of Alopoglossus lizard (Squamata, Gymnophthalmidae) from the tropical Andes, with a molecular phylogeny of the genus. ZooKeys 410: 105–120. doi: 10.3897/zookeys.410.7401 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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