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Andinosaura crypta (SÁNCHEZ-PACHECO, KIZIRIAN & SALES-NUNES, 2011)

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Common Names 
Proctoporus hypostictus BOULENGER 1902
Proctoporus hypostictus — HILLIS 1985: 109–126
Proctoporus hypostictus — KIZIRIAN & COLOMA 1991: 428 (part)
Proctoporus hypostictus — KIZIRIAN, 1995:72 (part)
Proctoporus hypostictus — KIZIRIAN 1996: 108 (part)
Proctoporus hypostictus — DOAN 2003: 372
Riama hyposticta — DOAN & CASTOE 2005: 409
Andinosaura crypta — SÁNCHEZ-PACHECO et al. 2017 
DistributionEcuador (western slopes of the Cordillera Occidental)

Type locality: Pilaló, Cotopaxi, Ecuador, 2320–2700 m elevation  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: KU 135103 
DiagnosisDiagnosis (genus Andinosaura) : All unambiguously optimized synapomorphies for this clade are from DNA sequences. Phenotypic synapomorphies are not known. Other characteristics include: (1) head scales usually smooth (slightly rugose in A. vieta and A. stellae); (2) frontoparietal and parietal scales paired; (3) interparietal, frontal and frontonasal scales single; (4) prefrontal scales usually absent (occasionally present in specimens of A. stellae); (5) lower eyelid divided into several scales; (6) loreal scale absent or present; (7) scale organs on labials present; (8) anteriormost supraocular and anteriormost superciliary scales unfused; (9) dorsal surface of the tongue covered in scale-like papillae; (10) nuchal scales smooth in most species (rugose in A. stellae and A. vieta); (11) dorsal body scales rectangular; smooth, keeled (low, rounded keel), striated (shallow furrows) or rugose; (12) ventral body scales smooth (rugose in A. stellae and A. vieta); (13) limbs pentadactyl; digits clawed; (14) femoral pores in males present, in females absent or present; and (15) hemipenial lobes large, distinct from hemipenial body.
Andinosaura differs from Riama by having hemipenial lobes large, distinct from hemipenial body. It differs from Oreosaurus by having a narrow band of differentiated granular lateral scales. [SÁNCHEZ-PACHECO et al. 2017]. 
CommentThis taxon was formerly referred to as Riama hyposticta, a rare species described on the basis of an adult male from northern Ecuador and here recorded from southwestern Colombia.

Type species: Riama crypta is the type species of the genus Andinosaura SÁNCHEZ-PACHECO et al. 2017. 
EtymologyAndinosaura (gender feminine), formed from the Spanish Andino (from the Andes) and the Latin Sauria (lizard), in reference to its distribution. The suffix is used commonly for gymnophthalmid genera. 
  • Sánchez-Pacheco, S. J., Torres-Carvajal, O., Aguirre-Peñafiel, V., Sales-Nunes, P. M., Verrastro, L., Rivas, G. A., Rodrigues, M. T., Grant, T. and Murphy, R. W. 2017. Phylogeny of Riama (Squamata: Gymnophthalmidae), impact of phenotypic evidence on molecular datasets, and the origin of the Sierra Nevada de Santa Marta endemic fauna. Cladistics, doi:10.1111/cla.12203 [print: 2018] - get paper here
  • Sánchez-Pacheco, Santiago J.; David A. Kizirian, and Pedro M. Sales-Nunes 2011. A New Species of Riama from Ecuador Previously Referred to as Riama hyposticta (Boulenger, 1902) (Squamata: Gymnophthalmidae). American Museum Novitates (3719): 1-15 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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