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Andinosaura kiziriani (SÁNCHEZ-PARECO, AGUIRRE-PENAFIEL & TORRES-CARVAJAL, 2012)

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymRiama aurea SÁNCHEZ-PARECO, AGUIRRE-PENAFIEL & TORRES-CARVAJAL 2012
Andinosaura kiziriani — SÁNCHEZ-PACHECO et al. 2017
Riama kiziriani — FELDMAN et al. 2015 
DistributionEcuador

Type locality: ECUADOR: Azuay: San Antonio, 1900 m elevation  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
Typesholotype: QCAZ 9667;
paratype: from El Chorro de Giron, Giron, 2546 m (QCAZ 9607) 
DiagnosisDiagnosis – Riama kiziriani can be distinguished from its congeners, except R. vespertina, by the presence of two anteriorly and one posteriorly positioned superciliary scales. it can be distinguished from R. vespertina by the following characteristics (condition for R. vespertina in parentheses): femoral pores per hind limb in males seven (4-5); and venter dark brown to black with small white spots or narrow lines on longitudinal sutures (ventral coloration in males cream – yellowish in life – with faint brown spots on central portion of scales).
in addition, among the other four species of Ri‐ ama currently known to occur in southern ecuador, R. kiziriani differs from R. anatoloros in having two postparietals (three in R. anatoloros), 20 longitudinal dorsal scale rows in males (22-27), and 32-34 transverse dorsal scale rows in males (36-42). it differs from R. stigmatoral in having three supraoculars (four in R. stigmatoral), seven femoral pores per hind limb in males (9-11), and six scales between medialmost femoral pores in males (0-2). it can be distinguished from R. petrorum by the second supraocular contacting the ciliaries (second, third and fourth, or first, second and third in contact with ciliaries in R. petrorum), by the number of femoral pores per hind limb in males, seven (4-5), and by the number of scales between medialmost femoral pores, six (eight); it further differs from R. petrorum in adult body size (maximum known svL in R. kiziriani is about 61 mm for males, versus 72 mm in R. petrorum). from R. au‐ rea it differs in having three supraoculars (four in R. aurea), seven femoral pores per hind limb in males (four) and venter dark brown to black with longitudinally arranged white stripes or spots on the scale sutures (venter cream with brown smears in scale centers gradually becoming more distinct ventrolaterally and on tail, forming nearly continuous longitudinal lines) (Sanchez-Pacheco et al. 2012: 263). 
Comment 
EtymologyNamed after David A. Kizirian, “our friend and colleague, in recognition of his contribution to the knowledge of Riama diversity”, and used as a noun in the genitive case. 
References
  • Sánchez-Pacheco, S. , Aguirre-Peñafiel, V. , Torres-Carvajal, O. 2012. Lizards of the genus Riama (Squamata: Gymnophthalmidae): the diversity in southern Ecuador revisited. South American J. Herp. 7 (3): 259-275 - get paper here
  • Sánchez-Pacheco, S. J., Torres-Carvajal, O., Aguirre-Peñafiel, V., Sales-Nunes, P. M., Verrastro, L., Rivas, G. A., Rodrigues, M. T., Grant, T. and Murphy, R. W. 2017. Phylogeny of Riama (Squamata: Gymnophthalmidae), impact of phenotypic evidence on molecular datasets, and the origin of the Sierra Nevada de Santa Marta endemic fauna. Cladistics, doi:10.1111/cla.12203 [print: 2018] - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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