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Anolis occultus WILLIAMS & RIVERO, 1965

IUCN Red List - Anolis occultus - Least Concern, LC
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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Puerto Rican twig anole, Limestone Anole, Dwarf Anole
Spanish: Largartijo enano 
SynonymAnolis occultus WILLIAMS & RIVERO 1965: 4
Anolis occultus — SCHWARTZ & HENDERSON 1991: 306
Anolis occultus — NICHOLSON et al. 2005
Deiroptyx occulta — NICHOLSON et al. 2012
Deiroptyx occulta — NICHOLSON et al. 2018 
DistributionPuerto Rico

Type locality: Road 143, midway beteen Cerro La Punta (1338 m) and Cerro Maravilla (1183 m), Puerto Rico.  
Reproductionoviparous 
TypesHolotype: MCZ 80303. 
Diagnosis 
CommentFor illustrations see Williams and Rivero, 1965; Thomas, 1965; Rivero, 1978; Gorman, 1981.

Species group: Deiroptyx occulta species group (fide NICHOLSON et al. 2012). 
References
  • Gorman, G.C. 1981. Anolis occultus, a small cryptic canopy lizard: are there pair bonds? Carib. J. Sci. 15 (3-4):29-31.
  • Huie, Jonathan M; Ivan Prates, Rayna C Bell, Kevin de Queiroz 2021. Convergent patterns of adaptive radiation between island and mainland Anolis lizards. Biological Journal of the Linnean Society, 2021;, blab072, - get paper here
  • Losos, J.B. 2007. Detective Work in the West Indies: Integrating Historical and Experimental Approaches to Study Island Lizard Evolution. BioScience 57 (7): 585-597 - get paper here
  • Nicholson, K. E., R. E. Glor, J. J. Kolbe, A. Larson, S. B. Hedges, and J. B. Losos 2005. Mainland colonization by island lizards. Journal of Biogeography 32: 929–938 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Patton, Austin H.; Luke J. Harmon, María del Rosario Castañeda, Hannah K. Frank, Colin M. Donihue, Anthony Herrel, and Jonathan B. Losos 2021. When adaptive radiations collide: Different evolutionary trajectories between and within island and mainland lizard clades. Proc. Natl. Acad. Sci. U.S.A., 10.1073/pnas.2024451118 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Rivero, J.A. 1978. Los anfibios y reptiles de Puerto Rico. M. Pareja Montana, 16, Barcelona, Espafia: x + 152 + 148pp.
  • Sanchez, Alejandro 2010. Observations on sleeping behavior of Anolis occultus and A. cuvieri in Puerto Rico. Anolis Newsletter VI: 174 - get paper here
  • Schwartz, A. 1974. A new species of primitive Anolis (Sauria, Iguanidae) from the Sierra de Baoruco, Hispaniola. Breviora (423): 1-19 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Thomas, R. 1965. Part Il. Field observations on Anolis occultus Williams and Rivero. Breviora (231):10-16. - get paper here
  • Webster, T. Preston 1969. Ecological observations on Anolis occultus Williams and Rivero (Sauria, Iguanidae). Breviora (312): 1-5 - get paper here
  • Williams, E. E. and J. A. Rivero. 1965. A new anole (Sauria, Iguanidae) from Puerto Rico. Part I. Description. Breviora (231):1-9. - get paper here
  • Yuan, M. L., M. H. Wake, and I. J. Wang. 2019. Phenotypic integration between claw and toepad traits promotes microhabitat specialization in the Anolis adaptive radiation. Evolution 73: 231–244 - get paper here
 
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