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Anolis campbelli (KÖHLER & SMITH, 2008)

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymNorops campbelli KÖHLER & SMITH 2008
Norops campbelli — NICHOLSON et al. 2012
Anolis campbelli — POE et al. 2017 
DistributionW Guatemala (Sierra de los Cuchumatanes, western Guatemala, from 1540 to 1660 m elevation).

Type locality: trail to Laguna Yolnabaj, Aldea Yalambojoch, Municipalidad Nentón, Departamento de Huehuetenango, Guatemala, 1540 m elevation, 16° 00’ 17’’ N, 91° 34’ 05’’ W.  
Reproductionoviparous 
TypesHolotype: UTA R-46038, an adult male, University of Texas at Arlington, collected 29 May 1998 by Manuel Acevedo and Marco V. Centeno. Original number MEA 1436. 
DiagnosisDiagnosis: A medium-sized species (snout– vent length [SVL] in largest specimen 51.3 mm) of the genus Norops (sensu Guyer and Savage, 1987, 1992) that belongs to the Norops schiedei species group (sensu Nieto-Montes de Oca, 2001). Norops campbelli differs from species of the N. schiedei group by the following characteristics (condition for N. campbelli in parentheses): Norops cobanensis: hemipenis small and unilobate (large and bilobate); scales comprising supraorbital semicircles usually distinctly larger than scales between semicircles (about subequal); smooth or only some faintly keeled midventral scales (most mid-ventral scales weakly, yet definitely keeled); some keels usually oblique or oriented transversely immediately anterior to prefrontal depression (oriented nearly longitudinally); scales in prefrontal depression mostly smooth (all scales in prefrontal depression strongly keeled); circumorbital row usually complete (rudimentary or almost absent). Norops cuprinus: prenasal distinct (prenasal fused with circumnasal); lorilabial row usually complete (usually incomplete). Norops cymbobs: para-vertebral scales slightly to moderately, albeit distinctly larger than scales in adjacent rows (paravertebral scales not larger than scales in adjacent rows); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); few or most preantebrachials and supra-antebrachials multicarinate (all unicarinate); prenasal distinct (prenasal fused with circumnasal). Norops duellmani: prenasal distinct (prenasal fused with circumnasal). Norops hobartsmithi: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal); scales in prefrontal depression mostly smooth (all scales in prefrontal depression strongly keeled); circumorbital row usually complete (rudimentary or almost absent); lorilabial row usually complete (usually incomplete). Norops johnmeyeri: postanal scales moderately enlarged in males (not enlarged); dewlap well developed in both sexes (not developed in females); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops matudai: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal); lorilabial row usually complete (usually incomplete). Norops milleri: paravertebral scales slightly to moderately, albeit distinctly, larger than scales in adjacent rows (paravertebral scales not larger than scales in adjacent rows); smooth or only some faintly keeled midventral scales (most mid-ventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops naufragus: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); middorsal scales between levels of axilla and groin 82–100 (68–69); few or most preantebrachials and supra-antebrachials multicarinate (all unicarinate). Norops parvicirculatus: scales comprising supraorbital semicir cles distinctly smaller than scales between semicircles (about subequal); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal); circumorbital row usually complete (rudimentary or almost absent). Norops pijolensis: middorsal scales between levels of axilla and groin 43–54 (68–69); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops polyrhachis: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled). Norops purpurgularis: horizontal loreal rows 7–9 (5–6); middorsal scales between levels of axilla and groin 52–62 (68– 69); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops schiedei: some keels usually oblique or oriented transversely immediately anterior to prefrontal depression and in depression (all head keels oriented nearly longitudinally). 
CommentSpecies group: Norops auratus Species Group (fide Nicholson et al. 2012). Not listed by Nicholson et al. 2018.

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet honors the foremost student of Guatemalan herpetology, Jonathan Campbell (1947-), American herpetologist, who also collected one of the type specimens and photographed the only two known specimens. See Murphy 1997 for biographical notes. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Köhler, Gunther and Eric N. Smith 2008. A New Species of Anole of the Norops schiedei group from Western Guatemala (Squamata: Polychrotidae). Herpetologica 64 (2): 216-223 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Murphy, J.B. 1997. A look at the life of Jonathan A. Campbell. Fauna 1 (1): 72-78
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Poe, Steven; Adrián Nieto-montes de Oca, Omar Torres-carvajal, Kevin De Queiroz, Julián A. Velasco, Brad Truett, Levi N. Gray, Mason J. Ryan, Gunther Köhler, Fernando Ayala-varela, Ian Latella 2017. A Phylogenetic, Biogeographic, and Taxonomic study of all Extant Species of Anolis (Squamata; Iguanidae). Syst Biol 66: 663–697 - get paper here
 
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