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Anolis argillaceus COPE, 1862

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Common NamesE: Cuban Dark Bark Anole, Bay Anole 
SynonymAnolis (Acantholis) argillaceus COPE 1862: 176
Anolis argillaceus — SCHWARTZ & HENDERSON 1991: 219
Anolis argillaceus — RODRÍGUEZ SCHETTINO 1999: 247
Anolis argillaceus — NICHOLSON et al. 2005
Anolis argillaceus — NICHOLSON et al. 2012
Anolis argillaceus — NICHOLSON et al. 2018 

Type locality: Monte Verde, Guantánamo Province, Cuba.  
TypesHolotype: lost, was USNM, lost fide Schwartz & Henderson 1988 
DiagnosisDESCRIPTION: Size small (SVL in males to 45 mm, in females to 36 mm); 2-3 rows of loreals; scales between supraorbitals 0; 2 scales between interparietal and supraorbital semicircles; 6 postrostrals; 7 postmentals; suboculars in contact with supralabials; supradigital scales smooth; body laterally compressed; limbs short; head scales smooth; dorsals smooth, small, flat; ventrals smooth, cycloid, in diagonal rows; tail laterally compressed, weakly verticillate; hemipenes very large, hemipenial swelling in tail extending to level of heel of adpressed foot; auricular opening round and without a fold. Dorsum with well-marked chestnut band between the fore- and hindlimbs; parallel to it (and separated by another grayish white band), another chestnut band, similar to the first; another much narrower and shorter parallel band from forelimbs onto tail; a third chestnut band,
joining with the second, on the side of face; median dorsal zone between neck and tail grayish beige; interparietal with a crescentic mark, oblique, dark chestnut, below it another smaller mark; a well-defined, dark chestnut interocular bar; remainder of head, lower eyelid, and throat yellow; 5 distinct bands on limbs; venter gray; dewlap peach to yellow or pale orange, at times with interior red to orange dots (Schwartz & Henderson 1991: 219). 
CommentIllustrations: Barbour and Ramsden, 1919; Garrido, 1975. Has been listed as synonym of A. loysiana by BOULENGER 1885.

Species group: Anolis loysianus species group (fide NICHOLSON et al. 2012). 
  • Alfonso, Yasel U.; Pierre Charruau, Gabriel Fajardo and Alberto R. Estrada. 2012. Interspecific communal oviposition and reproduction of three lizard species in Southeastern Cuba. Herpetology Notes 5: 73-77.
  • Barbour, T. and C. T. Ramsden. 1919. The herpetology of Cuba. Mem. Mus. Comp. Zool., Harvard Coll. 47: 69-213. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Cope, E.D. 1863. Contributions to Neotropical saurology. Proc. Acad. Nat. Sci. Philadelphia 14 [1862]: 176-188. - get paper here
  • Estrada, Alberto R. 1987. Anolis argillaceus (Sauria: Iguanidae): un nuevo caso de puestas comunales en anolis cubanos. Poeyana (353): 1-9
  • Garrido, O. H. 1988. Nueva especies para la ciencia de Anolis (Lacertilia: Iguanidae) de Cuba perteneciente al complejo argiliaceus. Donana, Acta Vert. 15: 45-57.
  • Garrido, O.H. 1975. Distribución y variación de Anolis argillaceus Cope (Lacertilia: Iguanidae) en Cuba. Poeyana (142):1-28.
  • Navarro, Nils P.; Fernandez, Alejandro V.; Garrido, Orlando H. 2001. Taxonomic reconsideracion of Anolis centralis litoralis and description of a new species of the argillaceus group (Sauria: Iguanidae) for Cuba. [in Spanish]. Solenodon 1:66-75
  • Nicholson, K. E., R. E. Glor, J. J. Kolbe, A. Larson, S. B. Hedges, and J. B. Losos 2005. Mainland colonization by island lizards. Journal of Biogeography 32: 929–938 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Rodríguez Schettino, Lourdes (ed.) 1999. The iguanid lizards of Cuba. University Press of Florida, 428 pp.
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Ruibal, R. 1964. An annotated checklist and key to the anoline lizards of Cuba. Bull. Mus. comp. Zool. Harvard 130 (8): 475-520. - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. and Ogren, L.H. 1956. A collection of reptiles and amphibians from Cuba, with the descriptionof two new forms. Herpetologica 12 (2): 91-110 - get paper here
  • Velasco, Julián A.; Fabricio Villalobos, José A. F. Diniz-Filho, Steven Poe and Oscar Flores-Villela 2020. Macroecology and macroevolution of body size in Anolis lizards. Ecography 00: 1–11 - get paper here
  • Williams, E. E.; Rand, A. S. 1977. Species recognition, dewlap function and faunal size. American Zoologist 17: 261-270 - get paper here
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