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Anolis marmoratus DUMÉRIL & BIBRON, 1837

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Higher TaxaDactyloidae, Iguania, Sauria (lizards) 
SubspeciesAnolis marmoratus alliaceus COPE 1864: 175
Anolis marmoratus caryae LAZELL 1964: 394
Anolis marmoratus girafus LAZELL 1964: 377
Anolis marmoratus inornatus LAZELL 1964: 386
Anolis marmoratus marmoratus DUMÉRIL & BIBRON 1837
Anolis marmoratus setosus LAZELL 1964: 380
Anolis marmoratus speciosus (GARMAN 1887) 
Common NamesGuadeloupean anole, Leopard Anole 
SynonymAnolis marmoratus DUMÉRIL & BIBRON 1837: 139
Anolis speciosus GARMAN 1888
Anolis marmoratus — UNDERWOOD in WILLIAMS et al. 1959
Ctenonotus marmoratus — SCHWARTZ & HENDERSON 1988: 113
Anolis marmoratus — SCHWARTZ & HENDERSON 1991: 297
Ctenonotus marmoratus — NICHOLSON et al. 2012

Anolis marmoratus alliaceus (COPE 1864)
Anolis alliaceus COPE 1864: 175
Anolis alliaceus — GÜNTHER 1888: 363
Anolis bimaculatus alliaceus — UNDERWOOD in WILLIAMS et al. 1959
Anolis marmoratus alliaceus — LAZELL 1964: 374
Anolis marmoratus alliaceus — SCHMIDT 1981
Ctenonotus marmoratus alliaceus — SCHWARTZ & HENDERSON 1988: 114
Anolis marmoratus alliaceus — HENKEL 2006
Ctenonotus marmoratus alliaceus — NICHOLSON et al. 2012

Anolis marmoratus caryae LAZELL 1964
Ctenonotus marmoratus caryae — SCHWARTZ & HENDERSON 1988
Anolis caryae — ROUGHGARDEN 1995
Ctenonotus marmoratus caryae — NICHOLSON et al. 2012

Anolis marmoratus girafus LAZELL 1964
Anolis marmoratus girafus — SCHMIDT 1981
Ctenonotus marmoratus girafus — SCHWARTZ & HENDERSON 1988
Ctenonotus marmoratus girafus — NICHOLSON et al. 2012

Anolis marmoratus inornatus LAZELL 1964
Anolis marmoratus inornatus — SCHMIDT 1981
Ctenonotus marmoratus inornatus — SCHWARTZ & HENDERSON 1988
Ctenonotus marmoratus inornatus — NICHOLSON et al. 2012

Anolis marmoratus setosus LAZELL 1964
Anolis marmoratus setosus — SCHMIDT 1981
Ctenonotus marmoratus setosus — SCHWARTZ & HENDERSON 1988
Ctenonotus marmoratus setosus — NICHOLSON et al. 2012

Anolis marmoratus speciosus (GARMAN 1887)
Anolis speciosus GARMAN 1887: 45
Anolis speciosus speciosus — UNDERWOOD in WILLIAMS et al. 1959
Anolis marmoratus speciosus — LAZELL 1964: 384
Ctenonotus marmoratus speciosus — SCHWARTZ & HENDERSON 1988: 115
Ctenonotus marmoratus speciosus — NICHOLSON et al. 2012 
DistributionLesser Antilles: Guadeloupe, Ile Desecheo Saintes, La Desirade,
French Guiana (introduced)

marmoratus: Guadeloupe: plain of Capesterre in southeastern Basse-Terre; intergrades with A. m. alliaceus, A. m. girafus, and A. m. speciosus.

alliaceus: central highlands in the Basse-Terre portion of Guadeloupe; intergrades with A. m. girafus, A. m. setosus, and A. m. speciosus.

caryae: Terre-de-Bas, Îles des Saintes; intergrades with A. m. terraealtae; introduced at Cayenne, Guiane Française.

girafus: leeward coast of Basse-Terre portion of Guadeloupe, from vicinity of Malendure and adjacent Ilets de Pigeon, south to Ballif; intergrades with A. m. setosus.

inornatus: northern Grande-Terre; Ilet Macou; intergrades with A. m. speciosus.

setosus northern coast of Basse-Terre portion of Guadeloupe from vicinity of Deshaies to Ste.-Rose; intergrades with A. m. speciosus.

speciosus: southwestern portion of Grande-Terre portion of Guadeloupe, onto isthmus connecting Grande-Terre and Basse-Terre; Ilet du Cochons, Ilet du Gosier, Ilet Christophe. Introduced to French Guiana (HOOGMOED, pers. comm.).

Type locality: Martinique (in error); revised to Capesterre on the Basse-Terreportion of Guadeloupe by Lazell, 1964. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesSyntypes: MNHN 794, MNHN 5491.
HOLOTYPE BMNH 1946.8.28.96 [alliaceus]
HOLOTYPE MCZ 70666 [caryae]
HOLOTYPE MCZ 71259 [girafus]
HOLOTYPE MCZ 71036 [inornatus]
HOLOTYPE MCZ 70813 [setosus]
LECTOTYPE MCZ 6172 [speciosus] 
CommentSubspecies: Anolis marmoratus ferreus, terraealtae, chrysops, desiradei, and kahouannensis are now considered a valid species.

Size: max snouth-vent-length (SVL) in mm (male/female): 82/54.
Size: max snouth-vent-length (SVL) in mm (male/female): 79/57 (alliaceus)
Size: max snouth-vent-length (SVL) in mm (male/female): 75/56 (girafus)
Size: max snouth-vent-length (SVL) in mm (male/female): 69/51 (setosus)
Size: max snouth-vent-length (SVL) in mm (male/female): 73/52 (speciosus)
Size: max snouth-vent-length (SVL) in mm (male/female): 65/50 (inornatus)
Size: max snouth-vent-length (SVL) in mm (male/female): 75/55 (caryae)

Species group: Ctenonotus bimaculatus species group (fide NICHOLSON et al. 2012). 
References
  • Barbour,T. 1915. Recent notes regarding West Indian reptiles and amphibians. Proc. Biol. Soc. Washington 28: 71-78 - get paper here
  • Breuil, H. 2002. Histoire Naturelle des Amphibiens et Reptiles de l’Archipel Guadeloupéen: Guadeloupe, Saint-Martin, Saint-Barthélemy. Patrimoines naturels 54: 339 pp.
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Garman, S. 1887. On West Indian reptiles. Iguanidae. Bull. Essex Inst. 19: 25-50.
  • Gorman, G.C. 1969. The zoogeography of Lesser Antillean Anolis lizards; an analysis based upon chromosomes and lactic dehydrogenases Bull. Mus. Comp. Zool. Harvard 138 (3): 53-80. - get paper here
  • Günther,A. 1888. Notes on reptiles and frogs from Dominica, West Indies. Ann. Mag. nat. Hist. (6) 2: 362-366 - get paper here
  • Hedges, S.B. & Conn, C.E. 2012. A new skink fauna from Caribbean islands (Squamata, Mabuyidae, Mabuyinae). Zootaxa 3288: 1–244 - get paper here
  • Hen kel, S. 2006. Haltung und Zucht von Anolis marmoratus alliaceus. Iguana-Rundschreiben 19 (2): 24-25
  • Hoogmoed, M.S., & Lescure, J. 1975. An annotated checklist of the lizards of French Guiana, mainly based on two recent collections Zool. Meded. (Leiden) 49(13): 141-172. - get paper here
  • Lazell,J.D. 1964. The anoles (Sauria: Iguanidae) of the Guadeloupéen archipelago.. Bull. Mus. comp. Zool. Harvard 131: 359-401 - get paper here
  • Lazell,J.D. 1972. The anoles (Sauria: Iguanidae) of the lesser Antilles. Bull. Mus. comp. Zool. Harvard 143 (1): 1-115 - get paper here
  • Lorvelec, Olivier; Pascal, Michel; Pavis, Claudie; Feldmann, Philippe. 2007. Amphibians and reptiles of the French West Indies: inventory, threats and conservation. Applied Herpetology 4 (2):131-161
  • Losos, J. B. 2009. Lizards in an Evolutionary Tree: Ecology and Adaptive Radiation of Anoles. Berkeley: University of California Press, 528 pp.
  • Malhotra, A. & Thorpe, R.S. 1999. Reptiles & Amphibians of the Eastern Caribbean. MacMillan, London & Oxford
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Roughgarden, J. 1995. Anolis Lizards of the Caribbean: Ecology, Evolution, and Plate Tectonics. OXFORD UNIVERSITY PRESS, 200 pp.
  • Rovatsos, Michail; Marie Altmanová, Martina Pokorná and Lukáš Kratochvíl 2014. CONSERVED SEX CHROMOSOMES ACROSS ADAPTIVELY RADIATED ANOLIS LIZARDS Evolution, DOI: 10.1111/evo.12357 - get paper here
  • Schmidt, J. 1981. Warum eigentlich immer Anolis carolinensis? Teil 4. Herpetofauna 3 (12): 6-8 - get paper here
  • Schmidt, Matthias 1996. Anolis marmoratus Duméril & Bibron, 1837 Reptilia 2 (8): 39-42 - get paper here
  • Schneider, C. J. 1996. Distinguishing between primary and secondary intergradation among morphologically differentiated populations of Anolis marmoratus Molecular Ecology 5: 239-249
  • Schwartz,A. & Henderson,R.W. 1988. West Indian Amphibian and Reptiles: A Checklist. Milwaukee Publ. Mus. Contr. Biol. Geol. No. 74: 264 pp.
  • Schwartz,A. & Henderson,R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Stenson, Andrew G.; Roger S.Thorpe and Anita Malhotra 2004. Evolutionary differentiation of bimaculatus group anoles based on analyses of mtDNA and microsatellite data. Molecular Phylogenetics and Evolution 32: 1 –10 - get paper here
  • Thorpe, R.S.; Jones, A.G.; Malhotra, A.; Surget-Groba, Y. 2008. Adaptive radiation in Lesser Antillean lizards: molecular phylogenetics and species recognition in the Lesser Antillean dwarf gecko complex, Sphaerodactylus fantasticus. Molecular Ecology 17(6):1489-1504
  • Williams, Ernest E.; Quesnel, Victor C.; Kenny, Julian S.; Underwood,G. 1959. The anoles of the Eastern Caribbean (Sauria, Iguanidae) Part. I. Preface; Part II. Two sibling species of anoles in Trinidad; Part. III. Revisionary notes. Bull. Mus. Comp. Zool. Harvard 121 (5): 185-226 + 1 plate - get paper here
 
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