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Anolis richardii DUMÉRIL & BIBRON, 1837

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Higher TaxaDactyloidae, Iguania, Sauria (lizards) 
Subspecies 
Common NamesGrenada Tree Anole 
SynonymAnolis Richardi DUMÉRIL & BIBRON 1837: 141
Anolis occipitalis GRAY 1840: 112
Anolis stenodactylus GRAY 1840: 114
Ptychonotus (Ctenodeira) richardii — FITZINGER 1843: 66
Anolis richardii — BOULENGER 1885: 37
Anolis trossulus GARMAN 1887: 38
Anolis trossulus — BARBOUR 1916
Anolis richardii richardii — UNDERWOOD in WILLIAMS et al. 1959
Anolis richardii — LAZELL 1972
Anolis richardi — SCHWARTZ & HENDERSON 1991: 324
Anolis richardii — NICHOLSON et al. 2005
Dactyloa richardii — NICHOLSON et al. 2012 
DistributionLesser Antilles: Grenadines, Grenada,Tobago, St. Vincent (SCHMIDT 1980)
Surinam (introduced according to HOOGMOED 1981: 281)

Type locality: “Tortola”. Revised to “Crown Point, Tobago” by Lazell 1972. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesHolotype: MNHN Ig. 53 (fide Underwood 1959); MCZ
Syntype: ANSP 23012; West Indies, Grenada; [trossulus]
Syntype: UMMZ, MCZ [trossulus] 
CommentSynonymy after Lazell 1972. According to Lazell (1972), “the species does not, and never did, occur on this small island [Tortola] in the British Virgins. I assume this to be a blunder, and suspect the real type locality as the somewhat similar sounding Tobago. I, therefore, revise the type locality to Crown Point, Tobago, where this species is abundant.”

Size: max snouth-vent-length (SVL) in mm (male/female):125/77.

Species groups: Dactyloa roquet species group (fide NICHOLSON et al. 2012). 
References
  • Barbour, T. 1916. Amphibians and reptiles of Tobago. Proc. Biol. Soc. Washington 29: 221-224 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Creer, Douglas A., Kevin de Queiroz, Todd R. Jackman, Jonathan B. Losos and Allan Larson 2001. Systematics of the Anolis roquet series of the southern Lesser Antillies. Journal of Herpetology 35 (3): 428-441 - get paper here
  • Daudin, Jacques; de Silva, Mark. 2007. An annotated checklist of the amphibians and terrestrial reptiles of the Grenadines with notes on their local natural history and conservation. Applied Herpetology 4 (2):163-175
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Fitzinger, L. 1843. Systema Reptilium, fasciculus primus, Amblyglossae. Braumüller et Seidel, Wien: 106 pp. - get paper here
  • Garman, S. 1887. On West Indian reptiles. Iguanidae. Bull. Essex Inst. 19: 25-50.
  • Gorman, G.C. 1969. The zoogeography of Lesser Antillean Anolis lizards; an analysis based upon chromosomes and lactic dehydrogenases Bull. Mus. Comp. Zool. Harvard 138 (3): 53-80. - get paper here
  • Gorman, George C. & Atkins, Leonard 1968. The relationships of the Anolis of the roquet species group (Sauria: Iguanidae). III. Comparative study of display behavior. Breviora (284): 1-31 - get paper here
  • Hite, Jessica L.; Gomez, Carlos A. Rodriguez; Larimer, Samantha C.; Diaz-Lameiro, Alondra M.; Powell 2008. Anoles of St. Vincent (Squamata: Polychrotidae): population densities and structural habitat use. Caribbean Journal of Science 44(1):102-115
  • Hoogmoed, M. S. 1981. Introduced species of reptiles in Surinam. Notes on the herpetofauna of Surinam VIII Amphibia-Reptilia 1: 277-285 - get paper here
  • KÖHLER, GUNTHER 2014. Characters of external morphology used in Anolis taxonomy—Definition of terms, advice on usage, and illustrated examples. Zootaxa 3774 (2): 201–257 - get paper here
  • Lazell,J.D. 1972. The anoles (Sauria: Iguanidae) of the lesser Antilles. Bull. Mus. comp. Zool. Harvard 143 (1): 1-115 - get paper here
  • Malhotra, A. & Thorpe, R.S. 1999. Reptiles & Amphibians of the Eastern Caribbean. MacMillan, London & Oxford
  • Mertens, R. 1972. Herpetofauna tobagana Stuttgarter Beitr. zur Naturkunde nr. 252 22 pp. - get paper here
  • Nicholson, K. E., R. E. Glor, J. J. Kolbe, A. Larson, S. B. Hedges, and J. B. Losos 2005. Mainland colonization by island lizards. Journal of Biogeography 32: 929–938 - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Roughgarden, J. 1995. Anolis Lizards of the Caribbean: Ecology, Evolution, and Plate Tectonics. OXFORD UNIVERSITY PRESS, 200 pp.
  • Schmidt, J. 1980. Warum eigentlich immer Anolis carolinensis? Teil 2. Herpetofauna 2 (5): 22-25 - get paper here
  • Schwartz,A. & Henderson,R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Simmons, P. M., B. T. Greene, K. E. Williamson, R. Powell and J. S. J. Parmerlee 2005. Ecological interactions within a lizard community on Grenada. Herpetologica 61 (2): 124-134 - get paper here
  • Williams, Ernest E.; Quesnel, Victor C.; Kenny, Julian S.; Underwood,G. 1959. The anoles of the Eastern Caribbean (Sauria, Iguanidae) Part. I. Preface; Part II. Two sibling species of anoles in Trinidad; Part. III. Revisionary notes. Bull. Mus. Comp. Zool. Harvard 121 (5): 185-226 + 1 plate - get paper here
 
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