Apathya cappadocica (WERNER, 1902)
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Apathya cappadocica
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| Higher Taxa | Lacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards) |
| Subspecies | Apathya cappadocica cappadocica (WERNER 1902) Apathya cappadocica muhtari (EISELT 1979) Apathya cappadocica urmiana (LANTZ & SUCHOW 1934) Apathya cappadocica wolteri (BIRD 1936) |
| Common Names | E: Anatolian Lizard |
| Synonym | Lacerta cappadocica WERNER 1902: 1086 Latastia cappadocica — WERNER 1908: 79 Apathya cappadocica — MEHELY 1909: 431 Apathya cappadocica — LANTZ & SUCHOW 1934 Archaeolacerta (Apathya) cappadocica — SINDACO et al. 2000 Lacerta cappadocica — BISCHOFF 2002 Apathya cappadocica — MAYER & ARRIBAS 2003 Lacerta cappadocica cappadocica— BISCHOFF 2007 Apathya cappadocica — ARNOLD et al. 2007 Lacerta (Apathya) cappadocica — SINDACO & JEREMČENKO 2008 Apathya cappadocica — ŠMÍD et al. 2014 Apathya cappadocica cappadocica — YOUSEFKHANI et al. 2019 Apathya cappadocica muhtari (EISELT 1979) Lacerta cappadocica muhtari EISELT 1979: 413 Lacerta cappadocica schmidtlerorum EISELT 1979: 415 Lacerta (Apathya) cappadocica schmidtlerorum — SINDACO & JEREMČENKO 2008 Lacerta (Apathya) cappadocica muhtari — SINDACO & JEREMČENKO 2008 Apathya cappadocica muhtari — AFRASIAB et al. 2013 Apathya cappadocica muhtari — YOUSEFKHANI et al. 2019 Apathya cappadocica urmiana (LANTZ & SUCHOW 1934) Apathya cappadocica urmiana LANTZ & SUCHOW 1934 Lacerta cappadocica urmiana — EISELT 1979: 417 Apathya cappadocica urmiana — SCHMIDT 1939 Lacerta (Apathya) cappadocica urmiana — SINDACO & JEREMČENKO 2008 Apathya cappadocica urmiana — AFRASIAB et al. 2013 Apathya cappadocica muhtari — BÖHME 2014: 146 Apathya cappadocica urmiana — MAYER 2015 Apathya cappadocica urmiana — YOUSEFKHANI et al. 2019 Apathya cappadocica wolteri (BIRD 1936) Apathya cappadocica wolteri BIRD 1936: 266 Lacerta cappadocica wolteri — EISELT 1979: 412 Lacerta (Apathya) cappadocica wolteri — BISCHOFF 2005 Lacerta (Apathya) cappadocica wolteri — SINDACO & JEREMČENKO 2008 Apathya cappadocica wolteri — YOUSEFKHANI et al. 2019 |
| Distribution | Turkey (Cilician Taurus, east of the Euphrat River, region around Diyarbakir and Viran_ehir, east Siirt--izre, Amanus Mts. and surroundings), NE Iraq, NW Iran, W Syria muhtari: Turkey (SE Anatolia), N Iraq, NW Iran; Type locality: Tatvan, (Turkey) about 1800 m elevation schmidtlerorum: between Euphrates and the Lake Van, Type locality: “Vil. Diyarbakir: 10 km S Diyarbakir, Abzweigung Ovabag, Felsental” [Turkey] urmiana: Turkey (SE Anatolia), N Iraq, NW Iran. Type locality: “Kherra (= Cherra,) Schlucht des Flusses Berdesur, Persien, ca. 20 km SW Rezaiyeh” ( 37°26′55.3′′N 44°54′05.3′′E, 1550 m elevation) wolteri: S Turkey, NW Syria; Type locality: “3000 ft. [elevation], ten miles west of Gaziantep; on limestone rocks in thick scrub country”. Type locality: Cilic Taurus, (Turkey) 2000 m (elevation). |
| Reproduction | oviparous |
| Types | Syntype: lost, formerly ZMH R04403 (formerly no. 2647), “lost” fide Eiselt 1979. Holotype: NMW; Paratypes in ZFMK, SZE, NMW, ZSM, ZMH, BMNH, FMNH and CAS [muhtari] Holotype: ZSM 252/1977, male, collected by J. J. & J. F. Schmidtler, 25.04. 1977, paratypes: ZSM, NMW, ZFMK [schmidtlerorum] Holotype: BMNH (BML) 1935.11.4.128 Holotype: ZISP = ZIN 12657b, male [urmiana] Holotype: BMNH (also given as NHM) 1935.11.4.128, collected 23.04.1935 [wolteri] |
| Diagnosis | Diagnosis (genus). Unique among Lacertini in having clear single keels on scales beneath toes, a transparent window in the lower eyelid consisting of several black-edged scales, and variable scaling on the side of the snout that may involve up to three postnasal and three loreal scales and sometimes an additional scale between the rostral and the nostril. Also possessing the following features found only in a minority of other Lacertini: 9–10 premaxillary teeth, often 7–8 posterior presacral vertebrae with short ribs, outer edge of parietal scale not reaching lateral border of parietal table, often five upper labial scales anterior to subocular, no masseteric scale, in some populations eight longitudinal rows of ventral scales; preanal scale often very small, bordered by two or more semicircles of smaller scales; often broad light dorsolateral stripes on dorsum; blue ocelli often present on flanks; bright colouring on underside absent, at least posteriorly, no blue spots on outer ventral scales, kidney expanded anteriorly and more than half its length in front of sacrum, hemipenis with traces of an armature and its retracted lobes coarsely folded. More widely distributed features among Lacertini include head and body markedly depressed and supraocular osteoderms fenestrated in adults, usual number of presacral vertebrae in males 26, inscriptional ribs usually absent, tail brightly coloured in hatchlings, hemipenial microornamentationn of hook-shaped spines (from Arnold et al. 2007: 36). Diagnosis: “Eine Archaeolazerte; unterscheidet sich leicht von der nächstverwandten Art, Lacerta danfordi GUNTHER, 1876, durch das aus mehreren großen und kleineren schwärzlich gerandeten, durchsichtigen Schuppen im unteren Augenlid gebildete „Fenster", durch das lange erste Supratemporale, durch das Fehlen des Massetericums und durch die einfach gekielten Subdigitallamellen.” (Eiselt 1979: 410) Diagnose (muhtari): Eine Lacerta cappadocica; unterscheidet sich von L. c. cappadocica durch das scharf abgesetzte, keineswegs verschwommene Rückenmuster, besonders durch die gerad- oder kerbrandigen Supraciliarstreifen, den weniger stark gefleckten Kopf, die durchschnittlich geringere Zahl der Dorsalia, Gularia, Schenkelporen und Infradigitallamellen, der geringeren Zerteilung der 1. Supraocularia und den höheren perzentuellen Anteil an Parietalia mit konkavem Außenrand.—Von L. c. wolteri trennt sie das fast völlige Fehlen einer Querstreifung des Schwanzes, der weniger stark gefleckte Pileus, das Zurücktreten von Schwarzfärbung, die durchschnittlich geringere Zahl der Dorsalia, Supraciliargranula, Gularia und Schenkelporen von L. c. schmidtlerorum durch die durchlaufende Rückenstreifung ohne Zerfall der Streifen in Fleckenreihen; von L. c. urmiana durch das ungeteilte Anale und die 6 Längsreihen von Ventralschildern. (Eiselt 1979: 413). Diagnose (schmidtlerorum): “Eine Lacerta cappadocica; unterscheidet sich von allen anderen Subspezies durch die völlige Auflösung aller Elemente heller Längsstreifung in Flecken- bzw. Tupfenreihen, die bei Männchen in ein schwarzes Netzwerk eingeschlossen sind, während sie bei Jungtieren und Weibchen aus der einheitlich schwarzen oder dunkelbraunen Grundfarbe hervorleuchten. — 6 Ventralia-Längsreihen, Anale ungeteilt, durchschnittlich 60—64,2—69 Rückenschuppen, 19—20,2—27 Schenkelporen.” (Eiselt 1979: 416). |
| Comment | Synonymy: A. c. urmiana is much more closely related to A. yassujica than to the other subspecies of A. cappadocica (Mayer 2015). Mayer 2015 also states that schmidtlerorum is very similar to wolteri, indicating that both may be synonyms, which was confirmed by Hosseinian Yousefkhani et al. 2019. Types: For comments on the types of this species see BAUER & GÜNTHER 1995. Type Species: Lacerta cappadocica Werner, 1902 is the type species of the genus Apathya MÉHELY 1907. Apathya is a highly variable genus, which parallels its high morphological variation. Distribution: See map in SMID et al. 2014 for distribution in Iran. Etymology (muhtari): “Muhtar (türkisch) = Gemeindevorsteher. — Ich widme diese neue Subspezies in altbewährter Freundschaft Herrn Universitäts- professor Dr. Muhtar BASOGLU, Chef des Instituts für zoologische Systematik an der ,,T. C. Ege Üniversitesi" in Bornova/Izmir. Als erster Türke hat sich Professor BASOGLU der wissenschaftlichen Herpetologie gewidmet und hat mit seinen Schülern in Bornova ein Zentrum herpetologischer Forschung aufgebaut.” (Eiselt 1979: 413) |
| Etymology | Named after the Cappadocia region in Central Anatolia. |
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