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Apographon orestes (HARVEY & MUÑOZ, 2004)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common Names 
SynonymTomodon orestes HARVEY & MUÑOZ 2004
Tomodon orestes — WALLACH et al. 2014: 731
Apographon orestes — TREVINE et al. 2022 
DistributionBolivia (Tarija), Argentina (Salta, Jujuy)

Type locality: close to the Río Erquis, Reserva de Sama, Méndez province, Tarija, Bolivia, 2754 m elevation (21° 28’ 56’’ S, 64° 50’ 32’’ W).  
TypesHolotype: CBF 2316 
DiagnosisDiagnosis (genus). The following combination of characters distinguishes Apographon from all the other genera of the tribe: smooth dorsal scales in 17 rows at midbody; supralabials 7; preoculars 2; 31–38 pairs of paravertebral black ocelli along the body; slightly dark oral mucosa or oral palate black; hemipenis with long lobes, covered with spinulate calyces and long spines (Figure S7 in Trevine et al. 2022). 
CommentType species: Tomodon orestes HARVEY & MUÑOZ 2004 is the type species of the genus Apographon TREVINE et al. 2022. 
EtymologyNamed after the Greek noun “orestes”, used in apposition, meaning an inhabitant of mountains.

The genus Apographon was named after the Greek “Apographos” (masculine noun, translated as “copy” or “transcript”), in reference to its resemblance with the genus Tomodon, where it was previously allocated. 
  • Akmentins, M.S. & Vaira, M. 2010. Reptilia, Squamata, Dipsadidae, Tomodon orestes Harvey and Muñoz, 2004: Distribution extension, new country record. Check List 6 (2):248-249 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Burgos Gallardo, Freddy; Stazzonelli Sadir, Juan Carlos; Baldo, Jorge 2014. Ampliación del rango de distribución de Tomodon orestes Harvey y Muñoz, 2004 (Serpentes: Dipsadidae) para el territorio argentino. Cuad. Herpetol. 28: - get paper here
  • Harvey, M.B. & Muñoz S., A. 2004. A new species of Tomodon (Serpentes: Colubridae) from high elevations in the Bolivian Andes. Herpetologica 60 (3): 364-372 - get paper here
  • Scrocchi, Gustavo J.; Stazzonelli, Juan Carlos; Cabrera, Paula 2019. Nuevas citas de Squamata (Gekkonidae, Phyllodactylidae y Dipsadidae) para la provincia de Tucumán, Argentina. Cuad. herpetol. 33 (2): - get paper here
  • Trevine, V. C., Grazziotin, F. G., Giraudo, A., Sallesbery‐Pinchera, N., Vianna, J. A., & Zaher, H. 2022. The systematics of Tachymenini (Serpentes, Dipsadidae): An updated classification based on molecular and morphological evidence. Zoologica Scripta - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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