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Apostolepis intermedia KOSLOWSKY, 1898

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesKoslowsky’s Blackhead, Mato Grosso Burrowing Snake 
SynonymApostolepis intermedia KOSLOWSKY 1898: 30
Apostolepis intermedia — DE LEMA 2002
Apostolepis intermedia — MARTINS & DE LEMA 2015
Apostolepis intermedia — WALLACH et al. 2014: 51 
DistributionBrazil (Mato Grosso), Paraguay

Type locality: Mato Grosso, Brazil Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesHolotype: lost according to Lema (1993) and Ferraro & Williams (2006). 
CommentKnown only from the holotype (apparently lost fide Harvey 1999) and one other specimen.

Diagnosis. Apostolepis intermedia can be distinguished from other Apostolepis by the following combination of characters: three infralabials in contact with first pair of anterior chinshields, five stripes on dorsal region of body (paravertebral stripes reduced, at least in adult specimens – see below), six supralabials, four maxillary teeth separated from pair of enlarged fangs by short diastema, snout projected, and 33–37 subcaudals [ALBUQUERQUE & DE LEMA 2012]. 
  • ALBUQUERQUE, NELSON RUFINO DE & THALES DE LEMA 2012. Description of the second known specimen of Apostolepis intermedia (Serpentes, Colubridae, Xenodontinae). Zootaxa 3325: 53–58 - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • Entiauspe-Neto, Omar Machado; Thales de Lema, Hugo Enrique Cabral Beconi 2014. Apostolepis intermedia Koslowsky, 1898 (Serpentes: Xenodontinae: Elapomorphini): First records for Paraguay. Check List 10 (3): 600-601 - get paper here
  • Ferraro, D.P. & Williams, J.D. 2006. Material tipo de la Colección de Herpetología del Museo de La Plata, Buenos Aires, Argentina. Cuadernos de Herpetologia, 19: 19–36
  • Koslowsky,J. 1898. Ofidios de Matto-Grosso (Brazil). Rev. Mus. La Plata 8: 25-34 - get paper here
  • Lema, T. de 2004. Nova espécie de Apostolepis COPE do estado de RONDÔNIA, Brasil (Serpentes, Elapomorphinae). Comunicações do Museu Ciência Tecnologia (PUCRS), Porto Alegre, 17 (2): 81-89
  • Lema, T. de & Renner, M.F. 2006. A new species of Apostolepis with striped pattern from Mato Grosso, Brasil (Serpentes, Elapomorphine). Ciência em movimento 8: 13-18
  • Lema, Thales de 2001. Fossorial snake genus Apostolepis from South America (Serpentes: Colubridae: Elapomorphinae). Cuadernos de herpetología 15 (1): 29-43.
  • Martins, L.A. & Lema, T. de 2015. Elapomorphini (Serpentes, Xenodontinae) do Brasil sudoeste. Neotropical Biology and Conservation 10(2): 93-102; doi: 10.4013/nbc.2015.102.05 - get paper here
  • Nogueira, Cristiano; Fausto Erritto Barbo and Hebert Ferrarezzi 2012. Redescription of Apostolepis albicollaris Lema, 2002, with a Key for the Species Groups of the Genus Apostolepis (Serpentes: Dipsadidae: Elapomorphini). South American J. Herp. 7 (3): 213-225 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
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