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Arrhyton dolichura WERNER, 1909

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Higher TaxaColubridae (Dipsadinae), Alsophiini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Havana Racerlet, Habana Island Racer 
SynonymArrhyton dolichurum WERNER 1909: 224
Arrhyton dolichura — SCHWARTZ & HENDERSON 1991: 584
Arrhyton dolichura — CROTHER 1999
Arrhyton dolichurum — ZAHER et al. 2009
Arrhyton dolichura — WALLACH et al. 2014: 56
Arrhyton dolichura — DÍAZ et al. 2021 

Type locality: La Habana, La Habana Province, Cuba.  
TypesHolotype: ZMH, now destroyed. 
DiagnosisDESCRIPTION: Maximum SVL 249 mm; ventrals 127-132 in males, 123-132 in females; subcaudals 101-127 in males, 104-127 in females; ventrals + subcaudals 232-254 in males, 229-252 in females; dorsal scale rows at midbody 17; supralabials 7; infralabials 8-9; preoculars 1/1; postoculars usually 2/2; loreal 1/1; anal divided; rostral slightly upturned; head neither conspicuously flattened nor broad. Dorsal pattern a series of 3 brown lines, the lateralmost lying on scale rows 3 and 4; dorsal scale rows 1 and 2 and lower portion of 3 (i.e., all rows ventral to lateral lines) concolor with venter; dorsal cephalic pattern a dark brown cap of uniform intensity, vividly outlined in white; white pigment involving snout and lateral margins of brown cap from prefrontals to parietals; dark loreal line from naris to eye, and thence posteriorly across temporals to form dark body stripes; rostral and first few supralabials flecked with dark brown on pale to white ground; minute but conspicuous pair of dots paramedially, 1 on each side of parietal suture; ventral color white to cream (Schwartz & Henderson 1991: 585). 
CommentFor illustrations see Grant, Smith, and Dalmau, 1959; Schwartz, 1965; Schwartz and Garrido, 1981.

Distribution: see map in Díaz et al. 2021: Fig. 5. 
  • DÍAZ, L. M., FONG G., A., SALAS, D., & HEDGES, S. B. 2021. A new semifossorial snake of the genus Arrhyton (Squamata: Dipsadidae) from eastern Cuba, with taxonomic comments on other species. Zootaxa, 5052 (3): 406-418 - get paper here
  • Grant, C.,Smith, H.M., and (Alayo) Dalmau, P. 1959. The status of snakes of the genus Arrhyton in Cuba. Herpetologica 15 (3): 129-33. - get paper here
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96 - get paper here
  • Schwartz, A. 1965. A review of the colubrid snake genus Arrhyton with a description of a new subspecies from southern Oriente Province, Cuba. Proc. Biol. Soc. Washington 78: 99-113. - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Schwartz, A. and O. H. Garrido. 1981. A review of the Cuban members of the genus Arrhyton (Reptilia, Serpentes, Colubridae). Annals of the Carnegie Museum 50: 207-230. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner,F. 1909. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. Jahrb. Hamb. wiss. Anst. suppl. 2 (Mitt. Naturhist. Mus. Hamb. 1908) 26: 205-247 - get paper here
  • Zaher, Hussam; Grazziotin, Felipe Gobbi; Cadle, John E.; Murphy, Robert W.; Moura-Leite, Julio Cesar de; Bonatto, Sandro L 2009. Molecular phylogeny of advanced snakes (Serpentes, Caenophidia) with an emphasis on South American Xenodontines: a revised classification and descriptions of new taxa. Pap. Avulsos Zool. (São Paulo) 49 (11): 115-153 - get paper here
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