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Asaccus griseonotus DIXON & ANDERSON, 1973

IUCN Red List - Asaccus griseonotus - Least Concern, LC
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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Gray-marked Gecko
E: Nasrullah’s leaf-toed gecko [nasrullahi] 
SynonymAsaccus griseonotus DIXON & ANDERSON 1973
Asaccus griseonotus — KLUGE 1993
Asaccus griseonotus — RÖSLER 2000: 60
Asaccus nasrullahi WERNER 2006
Asaccus nasrullahi — PAPENFUSS et al. 2010
Asaccus griseonotus — PAPENFUSS et al. 2010
Asaccus griseonotus — NASRABADI et al. 2017
Asaccus nasrullahi — NASRABADI et al. 2017 
DistributionNE Iraq (Palegawra Cave),
W Iran (Kermanshah, Lorestan)

Type locality: 38.5 mi from Shahabad, Kennanshah, Kermanshahan [Kerman Prov., ex errore], Iran).

nasrullahi: SW Iran (Lorestan); Type locality: Shah
Bazan, 12 km NNE of (km 347 on the railway), near the small affluent Ab-I-Khornos, 600 m elevation.  
Reproductionoviparous 
TypesHolotype: FMNH 170824 Paratypes: FMNH
Holotype: ZMUC R 3447, female. Apparently collected by E.
Kaiser of the Danish Scientific Investigations in Iran (Schmidt, 1955). [Asaccus nasrullahi] 
DiagnosisDiagnosis: Anderson 1999: 136.


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CommentDate of publication: Eiselt published his description about 1 month after DIXON & ANDERSON.

Distribution: See map in SMID et al. 2014 for distribution in Iran.

Synonymy: Kamali et al. 2024 synonymized Asaccus nasrullahi with A. griseonotus based on molecular data. Nazarov et al. 2024 think that the synonym of ingae with griseonotus may be premature but conceded that “further studies are required to clarify the taxonomic status of Phyllodactylus ingae”. However, they also considered Asaccus ingae (Eiselt, 1973) as a valid species, while A. granularis Torki 2010 should most likely be considered as its junior synonym.

Similar species: Asaccus nasrullahi has been previously misidentified as Ptyodactylus hasselquisti (WERNER, cited in RASTEGAR-PUYANI 2006).

NCBI taxonID: 886396 [nasrullahi]

Etymology: A. nasrullahi was named after “Nasrullah Rastegar-Pouyani, in recognition of his contribution to the knowledge of the herpetology of Iran, including the genus Asaccus. Moreover, "nasrullah" means "victory of God" which concept seems appropriate for the survival, discovery and rediscovery of this gecko species” [from WERNER 2006]. 
EtymologyFrom grise-, modern Latin griseus = gray + Greek notos, not- = back. 
References
  • Afrasiab, Saman R. and Sarbaz I. Mohamad 2009. A study on cave-dwelling geckos in Iraq, with the description of a new species from Saffine mountain (Reptilia: Gekkonidae). Zoology in the Middle East 47:49-56 - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Arnold E N; Gardner A S 1994. A review of the middle eastern leaf-toed geckoes (Gekkonidae: Asaccus) with descriptions of two new species from Oman. Fauna of Saudi Arabia 14: 424-441 - get paper here
  • Baier, F. 2005. Herpetologische Reiseeindrücke aus Sri Lanka. Elaphe 13 (4): 59-64
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Dixon, J.R. & Anderson,S.C. 1973. A new genus and species of gecko (Sauria: Gekkonidae) from Iran and Iraq. Bull. South. Calif. Acad. Sci. 72: 155-160 [1975] - get paper here
  • Gholamifard, A 2011. Endemism in the reptile fauna of Iran. Iranian Journal of Animal Biosystematics 7 (1) - get paper here
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • NASRABADI, REZA; NASRULLAH RASTEGAR-POUYANI, ESKANDAR RASTEGAR-POUYANI, AHMAD GHARZI 2017. A revised key to the lizards of Iran (Reptilia: Squamata: Lacertilia). Zootaxa 4227 (3): 431–443 - get paper here
  • Nazari-Serenjeh, Farzaneh and Farhang Torki 2008. Additional specimens of the gecko Asaccus nasrullahi Werner, 2006 (Reptilia: Phyllodactylidae), with notes on taxonomy and ecology. Zoology in the Middle East 44: 57-66 - get paper here
  • PAPENFUSS, THEODORE J.; TODD R. JACKMAN, AARON M. BAUER, BRYAN L. STUART, MICHAEL D. ROBINSON, and JAMES F. PARHAM 2010. Phylogenetic Relationships Among Species of Southwest Asian Leaf-toed Geckos (Asaccus). Proceedings of the California Academy of Sciences, 61 (13): 587-596 - get paper here
  • Parsa, H.; H. Oraie; A. Khosravani; N. Rastegar Pouyani 2009. Systematics and Distribution of the Iranian Plateau Leaf-toed Geckos of the Genus Asaccus (Sauria: Gekkonidae). Iranian Journal of Animal Biosystematics 5 (2) - get paper here
  • Rastegar-Pouyani, N., Nilson, G. & Faizi, H. 2006. A new Species of Asaccus (Sauria: Gekkonidae) from Kurdistan Province, western Iran. Hamadryad 30: 141-150. - get paper here
  • Rastegar-Pouyani, Nasrullah 2006. Systematics of the genus Asaccus (Sauria: Gekkonidae) on the Zagros Mountains, Iran. Herpetologica Bonnensis II: 117-119 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097 - get paper here
  • Tamar, Karin; Pelagia Mitsi; Salvador Carranza 2018. Cryptic diversity revealed in the leaf‐toed gecko Asaccus montanus (Squamata, Phyllodactylidae) from the Hajar Mountains of Arabia. J Zool Syst Evol Res. 2018 ;00:1–14 - get paper here
  • Torki F., Gharzi A., Nazari-Serenje F., Javanmardi S., Heidari N., Azizpourian A., Mahdavi S., and Mahdavi-Zarkhoni M. A. 2008. Geckos of the Genera Tropiocolotes and Asaccus in the Zagros Mountains, Iran. Gekko 5 (2): 31-43
  • TORKI, F. 2010. Beschreibung eines neuen Asaccus (Reptilia: Phyllodactylidae) aus der Provinz Lorestan, Iran. Sauria 32 (4): 3-16 - get paper here
  • Torki, F. 2011. Beschreibung eines neuen Asaccus (Sauria: Phyllodactylidae) aus dem Iran. Sauria 33 (1): 51–62 - get paper here
  • Torki, F.; Ahmadzadeh, F.; Ilgaz, C.; Avci, A. & Kumlutas, Y. 2011. Description of four new Asaccus Dixon and Anderson, 1973 (Reptilia: Phyllodactylidae) from Iran and Turkey. Amphibia-Reptilia 32 (2): 185-202 - get paper here
  • Torki, Farhang; Nasrin Heidari, Muhammad Sharif Khan 2010. A Morphological and Ecological Study of Asaccus nasrullahi Werner, 2006 (Reptilia: Phyllodactylidae), in Western Iran. Russ. J. Herpetol. 17 (3): 195-201 - get paper here
  • Werner, Y. L. 2006. Retraction of Ptyodactylus Goldfuss from the fauna of Iran and its replacement by a new species of Asaccus Dixon & Anderson (Reptilia: Sauria: Gekkonidae). Hamadryad 30: 135-140. - get paper here
 
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