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Asaccus elisae (WERNER, 1895)

IUCN Red List - Asaccus elisae - Least Concern, LC

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Higher TaxaPhyllodactylidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesWerner's Leaf-Toed Gecko 
SynonymPhyllodactylus elisae WERNER 1895
Phyllodactylus eugeniae NIKOLSKY 1907
Phyllodactylus elisae — SCHMIDT 1939
Phyllodactylus eliasae — HAAS [sic] 1952
Phyllodactylus ingae EISELT 1973
Phyllodactylus elisae — ARNOLD 1977: 100
Asaccus elisae — DIXON & ANDERSON 1973
Asacus [sic] elisae — BARAN & GRUBER 1982
Asaccus elisae — KLUGE 1993
Asaccus elisae — RÖSLER 1995: 98
Asaccus elisae — PAPENFUSS et al. 2010 
DistributionS Turkey (Birecik, SE Anatolia, Sanliurfa: HR 33: 223)
E Iraq, SW Iran, NE Syria (Niniveh)

Type locality: ruins of Niniveh (near Mosul)

eugeniae: Iran; Type locality: Dizful and Abu-Garia, affluent to Karun River, Iran. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesSyntypes: NMW (4 spec.); BMNH 95.3.2.3 (1946.8.24.39)
Syntypes ZIL 10261 [6+], 10262 [6+], 10263 [4], 10270[3], 10349) [Phyllodactylus eugeniae] 
CommentNot listed by Engelmann et al. (1993).

Synonymy: mostly after ARNOLD & GARDNER 1994.

Diagnosis: Easily distinguishable from other Asaccus species by the strongly keeled dorsal tubercles, the length of the tubercles being equal to the length of the oval ear opening, enlarged tubercles on upper arm, and whorls of keeled and pointed tubercles and scales on the tail. Tail length 55-70 mm, thinner and much longer than snout-vent length. Body slender, about 45-50 mm; dorsum with two series of dark spots on each side of vertebral line. Male with 4 wide black bands on the tail. Lamellae adhesive pads smaller than ear opening, claws extending somewhat the digit scansors.

Diagnosis: Relatively large (57 mm from snout to vent); two pairs of postmentals bordered by 18- 20 granules; scales on supraorbital region coarse, as are those of snout (11-16 between postnasal scales and orbit, 14-19 across snout at level of third upper labial);tubercles of dorsum, limbs and tail large, length of individual tubercle more than 64% of ear diameter; 8-14 longitudinal rows of enlarged dorsal tubercles; 10-12 large tubercles across rear of head between ears; 2-12 enlarged tubercles on upper forelimb above elbow; tail tubercles arranged in whorls, each whorl separated from other such tubercles by 2-3 granules; subtibial scales coarse; digital scansors not extending well beyond claws; phalanges in fourth toe reduced to four; cloacal tubercle small; tail tip laterally compressed; subcaudal series of expanded scales not reaching vent area anteriorly; tail color not sexually dimorphic, with a series of dark transverse bars that extend ventrally (Dixon and Anderson, 1973; Arnold and Gardner, 1994; Anderson, 1999; Rastegar-Pouyani et al., 2007) [from Parsa et al. 2009]

Type species: Phyllodactylus elisae WERNER 1895 is the type species of the genus Asaccus DIXON & ANDERSON 1973.

Diagnosis (genus): Digits with paired terminal scansors that lack lamellae, reduction of phalangeal formula of manus to 2.3.4.4.3, no transverse processes on autotomic caudal vertebrae except sometimes the first (ARNOLD 1977), no femoral pores, cloacal sacs or bones, only one egg laid at a time and left oviduct absent. The left oviduct is present in Phyllodactylus and in the apparently related Paroedura [ARNOLD & GARDNER 1994]. 
References
  • Afrasiab, Saman R. and Sarbaz I. Mohamad 2009. A study on cave-dwelling geckos in Iraq, with the description of a new species from Saffine mountain (Reptilia: Gekkonidae). Zoology in the Middle East 47:49-56 - get paper here
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Anderson,S.C. 1963. Amphibians and Reptiles from Iran. Proc. Cal. Acad. Sci. Ser. 4, 31(16): 417-498 - get paper here
  • Arnold E N; Gardner A S 1994. A review of the middle eastern leaf-toed geckoes (Gekkonidae: Asaccus) with descriptions of two new species from Oman. Fauna of Saudi Arabia 14: 424-441 - get paper here
  • Arnold, E. N. 1977. Little-known geckoes (Reptilia: Gekkonidae) from Arabia with descriptions of two new species from the Sultanate of Oman. The Scientific Results of the Oman Flora and Fauna Survey, 1975. Journal of Oman Studies, Special Report, (1):81—110 [1975]
  • Baran, Ibrahim; Yusuf Kumlutas; Cetin Ilgaz; Oguz Turkozan, and Aziz Avci 2003. ON Asaccus elisae (F. WERNER, 1895) (SAURIA: GEKKONIDAE) SPECIMENS COLLECTED FROM THE VICINITY OF HARRAN, SANLIURFA IN SOUTHEAST TURKEY. Russ. J. Herpetol. 10 (1): - get paper here
  • Dixon,J.R. & Anderson,S.C. 1973. A new genus and species of gecko (Sauria: Gekkonidae) from Iran and Iraq. Bull. South. Calif. Acad. Sci. 72: 155-160 [1975] - get paper here
  • Eiselt, J. 1973. Ein neuer Blattfinger-Gecko (Phyllodactylus, Sauria, Rept.) aus dem Iran und Bemerkungen zu Phyllodactylus elisae. Annalen des Naturhistorischen Museums in Wien, 77:173—179 - get paper here
  • Esterbauer, H. 1992. Asaccus elisae (Werner 1895) in Syrien. Sauria 14 (3): 31-34 - get paper here
  • Fathnia, B.; N. Rastegar Pouyani; M. Sampour; A.M. Bahrami; G. Jaafari 2009. The lizard fauna of Ilam province, Southwestern Iran. Iranian Journal of Animal Biosystematics 5 (2) - get paper here
  • Haas G. and Werner, Y. L. 1969. Lizards and snakes from Southwestern Asia, collected by Henry Field. Bull. Mus. Comp. Zool. Harvard 138: 327-406. - get paper here
  • LEPTIEN, R., T. KOWALSKI & H.J. ZILGER 1994. Der "Große Asaccus" aus den Omanbergen, Asaccus elisae (WERNER, 1895) s. lat. Sauria 16 (1): 21-25 - get paper here
  • Leviton,A.E.; Anderson,S.C.; Adler, K.; Minton,S.A. 1992. Handbook to Middle East Amphibians and Reptiles. SSAR, Oxford, Ohio (Contr. to Herpetol. No. 8), 1-252
  • Nikolsky, A. M. 1907. Reptiles et amphibiens recueillis (part.) M. N. A. Zarudny en Perse en 1903-1904 [in Russian and Latin]. Ann. Mus. Zool. Acad. Imp. Sci., St. Petersburg 10 [1905]: 260-301 - get paper here
  • PAPENFUSS, THEODORE J.; TODD R. JACKMAN, AARON M. BAUER, BRYAN L. STUART, MICHAEL D. ROBINSON, and JAMES F. PARHAM 2010. Phylogenetic Relationships Among Species of Southwest Asian Leaf-toed Geckos (Asaccus). Proceedings of the California Academy of Sciences, 61 (13): 587-596 - get paper here
  • Parsa, H.; H. Oraie; A. Khosravani; N. Rastegar Pouyani 2009. Systematics and Distribution of the Iranian Plateau Leaf-toed Geckos of the Genus Asaccus (Sauria: Gekkonidae). Iranian Journal of Animal Biosystematics 5 (2) - get paper here
  • Rastegar-Pouyani, N., Nilson, G. & Faizi, H. 2006. A new Species of Asaccus (Sauria: Gekkonidae) from Kurdistan Province, western Iran. Hamadryad 30: 141-150. - get paper here
  • Rastegar-Pouyani, Nasrullah 2006. Systematics of the genus Asaccus (Sauria: Gekkonidae) on the Zagros Mountains, Iran. Herpetologica Bonnensis II: 117-119 - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Rösler, Herbert 1995. Geckos der Welt - Alle Gattungen. Urania, Leipzig, 256 pp.
  • Schmidt,K.P. 1939. Reptiles and amphibians from Southwestern Asia. Publ. Field Mus. nat. Hist., zool. Ser., 24: 49-92 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, Roberto; Alberto Venchi, Giuseppe M. Carpaneto, Marco A. Bologna 2000. The Reptiles of Anatolia: a Checklist and Zoogeographical analysis. Biogeographia, 21-2000: 441-554.
  • ŠMÍD, JIŘÍ; JIŘÍ MORAVEC, PETR KODYM, LUKÁŠ KRATOCHVÍL, SEYYED SAEED HOSSEINIAN YOUSEFKHANI, ESKANDAR RASTEGAR-POUYANI & DANIEL FRYNTA 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa 3855 (1): 001–097
  • Tok, C.V.; Atatür, M.K.; Taskavak, E. 1997. Taxonomic status of Asaccus elisae (F. WERNER) from Birecik, Anatolia. Israel Journal of Zoology 43: 71-77
  • Tok, Cemal Varol; Mehmet Kutsay ATATÜR & Durmus CIHAN 2002. On an Asaccus elisae(F. Werner, 1895) (Sauria: Gekkonidae) Specimen Collected from the Vicinity of Nusaybin, Mardin. Turkish Journal of Zoology 26: 315-316 - get paper here
  • Torki F., Gharzi A., Nazari-Serenje F., Javanmardi S., Heidari N., Azizpourian A., Mahdavi S., and Mahdavi-Zarkhoni M. A. 2008. Geckos of the Genera Tropiocolotes and Asaccus in the Zagros Mountains, Iran. Gekko 5 (2): 31-43
  • TORKI, F. 2010. Beschreibung eines neuen Asaccus (Reptilia: Phyllodactylidae) aus der Provinz Lorestan, Iran. Sauria 32 (4): 3-16 - get paper here
  • Torki, F. 2011. Beschreibung eines neuen Asaccus (Sauria: Phyllodactylidae) aus dem Iran. Sauria 33 (1): 51–62 - get paper here
  • Torki, F.; Ahmadzadeh, F.; Ilgaz, C.; Avci, A. & Kumlutas, Y. 2011. Description of four new Asaccus Dixon and Anderson, 1973 (Reptilia: Phyllodactylidae) from Iran and Turkey. Amphibia-Reptilia 32 (2): 185-202 - get paper here
  • UGURTAS¸ Ismail HAKKI; HIKMET SAMI YILDIRIMHAN & Murat Sevinc 2007. Distribution of the Gekkonidae Species in Southeast Anatolia, Turkey, and New Localities. Turkish Journal of Zoology 31: 137-141 - get paper here
  • Vogel, G. 1989. Das Portrait: Asaccus elisae (WERNER). Sauria 11 (3): 1-2 - get paper here
  • Werner,F. 1895. Über eine Sammlung von Reptilien aus Persien, Mesopotamien und Arabien. Verh. Zool.-Bot. Ges. Wien 45: 13-22 - get paper here
 
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