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Aspidoscelis guttatus (WIEGMANN, 1834)

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Higher TaxaTeiidae, Teiinae, Gymnophthalmoidea, Sauria, Squamata (lizards)
SubspeciesAspidoscelis guttatus flavilineatus (DUELLMAN & WELLMAN 1960)
Aspidoscelis guttatus guttatus (WIEGMANN 1834)
Aspidoscelis guttatus immutabilis (COPE 1878) 
Common NamesE: Mexican Racerunner
S: Huico Ticuiliche Mexicano 
SynonymCnemidophorus guttatus WIEGMANN 1834: 29
Cnemidophorus unicolor COPE 1878: 93 (fide SMITH & TAYLOR 1950)
Cnemidophorus guttatus — GÜNTHER 1885: 28
Cnemidophorus guttatus unicolor — COPE 1892: 31
Cnemidophorus guttatus guttatus — GADOW 1906: 320
Cnemidophorus guttatus — BURT 1931: 66 (part.)
Cnemidophorus guttatus guttatus — SMITH & TAYLOR 1950: 179
Cnemidophorus guttatus guttatus — SMITH & BURGER 1950
Cnemidophorus guttatus guttatus — MASLIN & SECOY 1986
Cnemidophorus guttatus — LINER 1994
Cnemidophorus guttatus — KÖHLER 2000: 100
Aspidoscelis guttata — REEDER et al. 2002
Aspidoscelis guttata guttata — LINER & CASAS-ANDREU 2008
Aspidoscelis guttata — MATA-SILVA et al. 2015
Aspidoscelis guttatus — PYRON & BURBRINK 2013

Aspidoscelis guttatus flavilineatus (DUELLMAN & WELLMAN 1960)
Cnemidophorus guttatus flavilineatus DUELLMAN & WELLMAN 1960
Cnemidophorus guttatus flavilineatus — KLUGE 1984
Cnemidophorus guttatus flavilineatus — MASLIN & SECOY 1986
Aspidoscelis guttata flavilineata — REEDER et al. 2002
Aspidoscelis guttata flavilineata — LINER & CASAS-ANDREU 2008

Aspidoscelis guttatus immutabilis (COPE 1878)
Cnemidophorus immutabilis COPE 1878: 93
Cnemidophorus guttatus immutabilis COPE 1892: 31
Cnemidophorus microlepidopus COPE 1877: 92
Cnemidophorus guttatus microlepidopus — COPE 1892: 31
Cnemidophorus guttatus striatus GADOW 1903: 115
Cnemidophorus guttatus immutabilis — GADOW 1906: 320
Cnemidophorus immutabilis — WERNER 1910: 27
Cnemidophorus guttatus immutabilis — SMITH & TAYLOR 1950: 180
Cnemidophorus guttatus immutabilis — MASLIN & SECOY 1986
Aspidoscelis guttata immutabilis — REEDER et al. 2002
Cnemidophorus guttata immutabilis — CASTRO-FRANCO & BUSTOS-ZAGAL 2003
Aspidoscelis guttata immutabilis — LINER & CASAS-ANDREU 2008 
DistributionSE Mexico (Veracruz, Oaxaca, Guerrero through Chiapas)

Type locality: Mexico. Restricted to Veracruz, Veracruz, by SMITH & TAYLOR 1950.

flavilineatus: Mexico (Chiapas); Type locality: Mexico, Chiapas, Finca Orizaba, ca. 20 km SW Las Cruces, ca. 650 m elevation.

immutabilis: Mexico (Morelos)

striatus: Pacific slopes from Colima to Chiapas. Michoacán, Guerrero, Morelos. Type locality: “West Tehuantepec”, Oaxaca. Restricted to the city of Tehuantepec by SMITH & TAYLOR 1950.  
Reproductionoviparous 
TypesLectotype: ZMB 887 (designated by Burt, 1931:66, although given as 867, a lapsus calami; corrected by Duellman and Wellman, 1960:58); F. Deppe; 1824-1829
Holotype: USNM 30141; female; F. Sumichrast [immutabilis]
Holotype: UMMZ 119465; male; F. G. Thompson; 22 Dec. 1955 [flavilineatus]
Holotype: USNM 30385 [unicolor]
Holotype: USNM 30187 [Cnemidophorus microlepidopus] 
DiagnosisUnfortunately we had to temporarily remove additional information as this was scraped by multiple AI companies who sell that data to their customers. These details, e.g. detailed descriptions or comparisons (about 978 characters), are available for collaborators and contributors. Please contact us for details. 
CommentSynonymy after SMITH & TAYLOR 1950. Belongs to the Aspidoscelis deppii group (fide REEDER et al. 2002).

Types: Malnate 1971 listed 2 syntypes (ANSP 9579-80) for Cnemidophorus guttatus Hallowell 1855: 192. 
EtymologyNamed after Latin guttatus, provided with drops, spots or specks.[“...Color superne cineraceus, guttis albidis (in vivo animali pallide viridibus?), in series quatuor longitudinales dispositis,

A. g. flavilineatus was named after Latin flavus, yellow, golden, gold colored + Latin lineatus, lined. ["...Frorn the Latin, flavus, yellow, and lineatus, lined; used in reference to the broad yellow middorsal stripe in adults…"].

A. g. immutabilis was named after Latin immutabilis, unchangeable/unalterable. ["...This species attains to the largest size known in the genus, without losing its striped coloration, as do the other large forms…"]. 
References
  • Alvarez DEL TORO, M. 1982. Los Reptiles de Chiapas. 3rd ed. México: Tuxtla Gutiérrez, 248 pp.
  • Bauer, A.M. and R. Günther 1994. An annotated type catalogue of the teiid and microteiid lizards in the Zoological Museum, Berlin (Reptilia: Squamata: Teiidae and Gymnophthalmidae). Mitt. Zool. Mus. Berlin 70:267-280 - get paper here
  • BELLO-SÁNCHEZ, EDGAR AHMED; JORGE E. MORALES-MÁVIL & ORLANDO R. VIVANCO-MONTANÉ. 2021. ASPIDOSCELIS GUTTATUS (Mexican Whiptail Lizard). PREDATION. Herpetological Review 52 (3): 644. - get paper here
  • Burt, C. E. 1931. A study of the teiid lizards of the genus Cnemidophorus with special reference to their phylogenetic relationships. Bull. US Natl. Mus. 154: 286 pp. - get paper here
  • Casas-Andreu, G., F.R. Méndez-De la Cruz and X. Aguilar-Miguel. 2004. Anfibios y Reptiles; pp. 375–390, in A.J.M. García-Mendoza, J. Ordoñez and M. Briones-Salas (ed.). Biodiversidad de Oaxaca. Instituto de Biología, UNAM-Fondo Oaxaqueño para la Conservación de la Naturaleza-World Wildlife Fund, México, D. F.
  • Castro Franco, R. & M. G. Bustos-Zagal 2003. Lagartijas de Morelos, México: distribución, hábitat y conservación. Acta Zoológica Mexicana (n.s.) 88:123-142 - get paper here
  • Castro-Franco, Rubén, María Guasalupe Bustos-Zagal 1994. List of reptiles of Morelos, Mexico, and their distribution in relation to vegetation types. Southwestern Naturalist 39 (2): 171-175 - get paper here
  • Cope, E.D. 1878. Tenth contribution to the herpetology of Tropical America. Proc. Amer. Philos. Soc. 17: 85-98 [1877] - get paper here
  • Cope, E.D. 1892. A synopsis of the species of the teiid genus Cnemidophorus. Trans. Am. Phil. Soc. (ser. 2) 17 (1): 27-52 [1893] - get paper here
  • Duellman, W. E. & WELLMAN, J. 1960. A systematic study of the lizards of the deppei group (genus Cnemidophorus) in México and Guatemala. Misc. Publ. Mus. Zool. Univ. Michigan (No. 111): 1-81 - get paper here
  • Gadow,H. 1903. nan Proc. Roy. Soc. London 72: 115
  • Gadow,H. 1906. A contribution to the study of evolution based upon the Mexican species of Cnemidophorus. Proc. Zool. Soc. London 1906: 277-375 (+ maps) - get paper here
  • Günther, A. C. L. G. 1885. Reptilia and Batrachia. Biologia Centrali-Américana. Taylor, & Francis, London, 326 pp. [published in parts from 1885-1902; reprint by the SSAR 1987] - get paper here
  • Johnson, Jerry D.; Vicente Mata-Silva, Elí García Padilla, and Larry David Wilson 2015. The Herpetofauna of Chiapas, Mexico: composition, distribution, and conservation. Mesoamerican Herpetology 2 (3): 272–329. - get paper here
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • Lemos-Espinal JA, Smith GR 2020. A conservation checklist of the herpetofauna of Morelos, with comparisons with adjoining states. ZooKeys 941: 121-144 - get paper here
  • Lemos-Espinal JA, Smith GR, Pierce LJS, Painter CW 2020. The amphibians and reptiles of Colima, Mexico, with a summary of their conservation status. ZooKeys 927: 99-125 - get paper here
  • Malnate, Edmond V. 1971. A catalogue of primary types in the Herpetological Collections of the Academy of Natural Sciences, Philadelphia (ANSP). Proccedings of the Academy of Natural Sciences of Philadelphia 123 (9): 345-375 - get paper here
  • Maslin, T. & Secoy, D.M. 1986. A checklist of the lizard genus Cnemidophorus (Teiidae). Contr. Zool. Univ. Colorado Mus. 1: 1-60
  • Mata-González, Stephanye, Vicente Mata-Silva, Dominic L. DeSantis, Elí García-Padilla and Larry David Wilson. 2016. Aspidoscelis guttata (Wiegmann, 1834). Opportunistic water acquisition. Mesoamerican Herpetology 3 (2): 483-484 - get paper here
  • Mata-Silva, Vicente, Jerry D. Johnson, Larry David Wilson and Elí García-Padilla. 2015. The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology 2 (1): 6–62 - get paper here
  • Palacios-Aguilar, Ricardo & OSCAR FLORES-VILLELA 2018. An updated checklist of the herpetofauna from Guerrero, Mexico. Zootaxa 4422 (1): 1-24 - get paper here
  • Peterson, A.T. et al. 2004. A preliminary biological survey of Cerro Piedra Larga, Oaxaca, Mexico: Birds, mammals, reptiles, amphibians, and plants. Anales del Instituto de Biología, Universidad Nacional Autónoma de México, Serie Zoología 75(2): 439-466
  • Pyron, R. Alexander; Frank T. Burbrink 2013. Early origin of viviparity and multiple reversions to oviparity in squamate reptiles. Ecology Letters 17 (1): 13–21 (published online 2013, in print 2014), DOI: 10.1111/ele.12168 - get paper here
  • REEDER, T.W.; CHARLES J. COLE AND HERBERT C. DESSAUER 2002. Phylogenetic Relationships of Whiptail Lizards of the Genus Cnemidophorus (Squamata: Teiidae): A Test of Monophyly, Reevaluation of Karyotypic Evolution, and Review of Hybrid Origins. American Museum Novitates 3365: 1-64 - get paper here
  • Smith, H. M., & Burger, W.L. 1950. Herpetological results of the University of Illinois field expedition, Spring 1949. III. Sauria. Transactions of the Kansas Academy of Science 53 (2): 165-175 - get paper here
  • Torres-Hernández, LA, Ramírez-Bautista A, Cruz-Elizalde R, Hernández-Salinas U, Berriozabal-Islas C, DeSantis DL, Johnson JD, Rocha A, García-Padilla E, Mata-Silva V, Fucsko LA, and Wilson LD. 2021. The herpetofauna of Veracruz, Mexico: composition, distribution, and conservation status. Amphibian & Reptile Conservation 15(2) [General Section]: 72–155 - get paper here
  • Walker, J M; Corneil, J P; Kartje, M E; Montgomery, C E & Shandersky, L; 2019. Aspidoscelis guttatus (Mexican Racerunner) Juvenile characteristics. Herpetological Review 50 (2): 364-365 - get paper here
  • Walker, J M; Kartje, M E; Corneil, J P & Montgomery, C E; 2019. Aspidoscelis guttatus (Mexican Racerunner) and Sceloporus siniferus (Long-tailed Spiny Lizard) Opportunistic foraging. Herpetological Review 50 (3): 568-569 - get paper here
  • Werner,F. 1910. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. ii. Eidechsen. Jahrb. Hamburg. Wiss. Anst., vol. 27 (1909), suppl. no. 2, 1910, pp. 1-46; reprinted: 1910, Mitteil. Naturhist. Mus. Hamburg, vol. 27: 205-) - get paper here
  • Wiegmann,A.F.A. 1834. Herpetologia Mexicana, seu descriptio amphibiorum novae hispaniae , quae itineribus comitis de Sack, Ferdinandi Deppe et Chr. Guil. Schiede im Museum Zoologicum Berolinense Pervenerunt. Pars prima, saurorum species. Berlin, Lüderitz, iv + 54 pp. - get paper here
 
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