Asthenodipsas borneensis QUAH, GRISMER, LIM, ANUAR & CHAN, 2020
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Asthenodipsas borneensis
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| Higher Taxa | Pareidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) |
| Subspecies | |
| Common Names | E: Bornean Dark-necked Slug Snake |
| Synonym | Asthenodipsas borneensis QUAH, GRISMER, LIM, ANUAR & CHAN 2020 Asthenodipsas malaccanus — CHAN-ARD et al. 2015: 159 (part) Asthenodipsas malaccanus — CHARLTON 2020: 86–87 (part) Asthenodipsas malaccanus — DAS 2006: 007 (part) Asthenodipsas malaccanus — DAS 2010: 161 & 344–345 (part) Asthenodipsas malaccanus — DAS 2012 & 2018: 131 (part) Asthenodipsas malaccanus — ISKANDAR & COLIJN 2001: 111 (part) Asthenodipsas malaccanus — MARLON 2014: 27–28 (part) Asthenodipsas malaccanus — STUEBING et al. 2014: 87 (part) Asthenodipsas malaccanus — TEYNIÉ et al. 2010: 39 (part) Asthenodipsas malaccanus — WALLACH et al. 2014: 59 (part) Amblycephalus malaccanus — BOULENGER 1892: 507 (part) Amblycephalus malaccanus — BOULENGER 1896: 442–443 (part) Amblycephalus malaccanus — BOULENGER 1912: 209–210 (part) Amblycephalus malaccanus — FLOWER 1896: 896 (part) Amblycephalus malaccanus — FLOWER 1899: 607 & 694 (part) Amblycephalus malaccanus — DE ROOIJ 1917: 267–277 (part) Internatus malaccanus — DAVID & VOGEL 1996: 137–138 (part) Internatus malaccanus — MALKMUS et al. 2002: 343 Pareas malaccanus — DE HAAS 1950: 529 (part) Pareas malaccanus — HAILE 1958: 766 (part) Pareas malaccanus — GRANDISON 1972: 92 (part) Pareas malaccanus — MARLON 2014: 27–28 (part) Pareas malaccanus — STUEBING 1991: 331 (part) Pareas malaccanus — STUEBING & INGER 1999: 88 (part) Pareas malaccanus — TAYLOR 1965: 686 (part) Pareas malaccanus — WELCH 1988: 90 (part) Pareas carinatus — DAS 2006: 45 Pareas carinatus — DAS 2007: 161 Asthenodipsas (Asthenodipsas) borneensis — POYARKOV et al. 2022 |
| Distribution | East Malaysia (Borneo: Sabah) Type locality: Mendolong camp watershed, Sipitang District, Sabah, East Malaysia (4°54’N 115°45’E, 748 m elevation). |
| Reproduction | |
| Types | Holotype. FMNH 243935, Adult female, collected by Robert F. Inger and Tan Fu Lian on 25 August 1990. Paratypes. Adult male FMNH 243936 collected by Robert F. Inger and Tan Fu Lian on 9 September 1990 from Purulon camp, Tenom District, Sabah, East Malaysia (5°13’N 115°57’E, 516 m a.s.l.). Adult male FMNH 249751 collected by Robert F. Inger on 6 December 1991 from Poring station, Mount Kinabalu Park, Ranau District, Sabah, East Malaysia (6°03’N 116°42’E, 606 m a.s.l.). Adult female FMNH 71590 collected by T. H. Harrisson in May 1952 from Pa Brayong, Lawas District, Sarawak, East Malaysia (estimated: 4.45°N 115.51667°E, 1,077m a.s.l.). |
| Diagnosis | Diagnosis: Asthenodipsas borneensis sp. nov. is diagnosed as member of the genus Asthenodipsas by its pos- session of smooth dorsals, absence of preoculars and suboculars, having supralabials in contact with the eyes, single anterior inframaxillary and two pairs of posterior inframaxillaries (Grossmann & Tillack 2003). It can be separated from Aplopeltura boa (Boie) by its higher number of mid-dorsal scale rows (15 vs. 13) and subcaudals (divided vs. undivided) (de Rooij 1917; Grossmann & Tillack 2003; Stuebing et al. 2014), and can be further differentiated from members of the genus Pareas by its possession of preocular and subocular scales (absent vs. present), supralabials in contact with orbit (3rd & 4th contact orbit vs. no supralabials in contact with orbit) and anterior single inframaxil- lary (present vs. absent) (Grossmann & Tillack 2003). Asthenodipsas borneensis sp. nov. can be differentiated from its congeners by the following combination of characters: a maximum SVL of 441 mm; 15/15/15 dorsal scale rows; 166–179 ventrals; 35–48 subcaudal scales; two postoculars; 2(+0–1)+(1–3) temporals; seven or eight supralabials, 3rd & 4th touching the eye; 4–7 infralabials, 2nd or 3rd pair in contact; a sharp vertebral keel; dorsum of adults beige to orange-brown with a dark-brown to black patch on the neck followed by 23–40 irregularly-shaped, rhomboidal dark-brown bands that extend the length of body and tail and onto lateral edges of the ventral scales to form spots, but not encircling body; a narrow, light-coloured vertebral stripe starting from the nape or after the dark patch on the neck; throat and ventrals beige to yellow and mottled with small, dark spots except for neck region which is dark- brown to black, sometimes with a faint, light-coloured median stripe; head whitish to dark-grey with dark speckling on the snout, crown and labials; and iris reddish-brown and pupils black (Table 1,2 & 7; Fig. 2 & 3 in Quah et al. 2020). |
| Comment | Activity: nocturnal Diet: snails Habitat: lowland rainforests and montane regions up to elevations of 1000 m. During the day it reportedly shelters beneath leaf litter |
| Etymology | The specific epithet borneensis is in reference to its restriction to the island of Borneo. The suffix ensis is a Latin derivation meaning “from” or “inhabiting.” It renders the specific epithet an adjective that must be in grammatical accord with the gender of Asthenodipsas that is feminine (Frank Tillack in litt. 2019). |
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