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Atractus carrioni PARKER, 1930

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesParker's Ground Snake 
SynonymAtractus carrioni PARKER 1930: 208
Atractus carrioni — PETERS 1960
Atractus carrioni — PASSOS & ARREDONDO 2009
Atractus carrioni — WALLACH et al. 2014: 70 
DistributionEcuador (Loja)

Type locality: Loja, Ecuador, 2200 m elevation.  
Reproductionoviparous 
TypesHolotype: BMNH 1929.10.30.1, female 
DiagnosisDIAGNOSIS: An Atractus unique within the genus in having no loreal shield. Readily distinguished by the following combination of characteristics: (1) 15 scale rows; (2) no loreal, although a small preocular sometimes present; (3) maxillary teeth eight or nine; (4) ventrals in males, 145-149 (146) in females, 152-159 (155); and (5) dorsal coloration essentially uniform dark gray or brown [from SAVAGE 1960].

Diagnosis. Atractus carrioni can be distinguished from all congeners by the following combination of characters: (1) 15 smooth dorsal scale rows; (2) single postocular; (3) loreal absent; (4) temporals usually 1+1; (5) usually six supralabials, third and fourth contacting orbit; (6) usually six infralabials, first four contacting chinshields; (7) eight to ten maxillary teeth; (8) usually three gular scale rows; (9) usually four pre-ventrals; (10) 143–161 ventrals in females, 136–151 in males; (11) 18–28 subcaudals in females, 25–33 in males; (12) dorsum of body uniformly gray to grayish brown; (13) venter usually brown to grayish brown, occasionally brown pigment restricted to mid portion of ventral scales in which the lateral areas are cream; (14) moderate body size, females reaching 495 mm SVL, males 350 mm; (15) small to moderate tail in females (8.0–14.7% of SVL), moderate to long in males (9.9–17.2% of SVL); (16) hemipenis moderately bilobed, semicapitate and semicalyculate [Passos 2013: 111]. 
CommentSimilar species: A. roulei. The two species differ primarily in the presence in roulei and absence in carrioni of the loreal shield. 
EtymologyNamed after Professor Clodoveo Carrión Mora (1883-1957), and Ecuadorian natural scientist and paleontologist. After studying in Manchester and London, he returned to Ecuador he became Professor of Natural Sciences at the Colegio Bernardo Valdevisio. He collected the holotypes of the species. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • CISNEROS-HEREDIA, D. F. 2005. Report of molluscivory in Atractus carrioni PARKER, 1930 [Short Note]. Herpetozoa 18: 185- 186 - get paper here
  • Parker,H.W. 1930. A new colubrine snake from Ecuador. Ann. Mag. nat. Hist. (10) 5: 207-209
  • Passos, P. & Arredondo, J.C. 2009. Rediscovery and redescription of the Andean earth-snake Atractus wagleri (Reptilia: Serpentes: Colubridae). Zootaxa 1969: 59–68 - get paper here
  • Passos, Paulo; Lourdes Y. Echevarría, and Pablo J. Venegas 2013. Morphological Variation of Atractus carrioni (Serpentes: Dipsadidae). South American J. Herp. Aug 2013, Vol. 8, No. 2: 109-120. - get paper here
  • Savage,J.M. 1960. A revision of the Ecuadorian snakes of the colubrid genus Atractus. Misc. Publ. Zool. Univ. Michigan 112: 1-86 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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