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Atractus carrioni PARKER, 1930

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesParker's Ground Snake 
SynonymAtractus carrioni PARKER 1930: 208
Atractus carrioni — PETERS 1960
Atractus carrioni — PASSOS & ARREDONDO 2009
Atractus carrioni — WALLACH et al. 2014: 70 
DistributionEcuador (Loja)

Type locality: Loja, Ecuador, 2200 m elevation. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: BMNH 1929.10.30.1, female 
CommentDIAGNOSIS: An Atractus unique within the genus in having no loreal shield. Readily distinguished by the following combination of characteristics: (1) 15 scale rows; (2) no loreal, although a small preocular sometimes present; (3) maxillary teeth eight or nine; (4) ventrals in males, 145-149 (146) in females, 152-159 (155); and (5) dorsal coloration essentially uniform dark gray or brown [from SAVAGE 1960].

Diagnosis. Atractus carrioni can be distinguished from all congeners by the following combination of characters: (1) 15 smooth dorsal scale rows; (2) single postocular; (3) loreal absent; (4) temporals usually 1+1; (5) usually six supralabials, third and fourth contacting orbit; (6) usually six infralabials, first four contacting chinshields; (7) eight to ten maxillary teeth; (8) usually three gular scale rows; (9) usually four pre-ventrals; (10) 143–161 ventrals in females, 136–151 in males; (11) 18–28 subcaudals in females, 25–33 in males; (12) dorsum of body uniformly gray to grayish brown; (13) venter usually brown to grayish brown, occasionally brown pigment restricted to mid portion of ventral scales in which the lateral areas are cream; (14) moderate body size, females reaching 495 mm SVL, males 350 mm; (15) small to moderate tail in females (8.0–14.7% of SVL), moderate to long in males (9.9–17.2% of SVL); (16) hemipenis moderately bilobed, semicapitate and semicalyculate [Passos 2013: 111].

Similar species: A. roulei. The two species differ primarily in the presence in roulei and absence in carrioni of the loreal shield. 
References
  • CISNEROS-HEREDIA, D. F. 2005. Report of molluscivory in Atractus carrioni PARKER, 1930 [Short Note]. Herpetozoa 18: 185- 186 - get paper here
  • Parker,H.W. 1930. A new colubrine snake from Ecuador. Ann. Mag. nat. Hist. (10) 5: 207-209
  • Passos, P. & Arredondo, J.C. 2009. Rediscovery and redescription of the Andean earth-snake Atractus wagleri (Reptilia: Serpentes: Colubridae). Zootaxa 1969: 59–68 - get paper here
  • Passos, Paulo; Lourdes Y. Echevarría, and Pablo J. Venegas 2013. Morphological Variation of Atractus carrioni (Serpentes: Dipsadidae). South American J. Herp. Aug 2013, Vol. 8, No. 2: 109-120. - get paper here
  • Savage,J.M. 1960. A revision of the Ecuadorian snakes of the colubrid genus Atractus. Misc. Publ. Zool. Univ. Michigan 112: 1-86 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
 
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