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Bachia marcelae DONOSO-BARROS & GARRIDO, 1964

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Higher TaxaGymnophthalmidae (Cercosaurinae, Bachiinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common NamesE: Marcela’s Bachia, Marcela’s Worm lizard 
SynonymBachia marcelae DONOSO-BARROS & GARRIDO 1964: 3
Bachia lineata marcelae — DONOSO-BARROS 1968: 118
Bachia heteropa marcelae – DIXON 1973: 33
Bachia heteropa marcelae — JOHN et al. 2012
Bachia marcelae — MURPHY et al. 2019 
DistributionVenezuela

Type locality: “Bosque tropical La Luz, Edo. Barinas, Venezuela.”, given as La Luz, Edo. Barinas, Venezuela (8° 23′ N, 69° 49′ W) by Murphy et al. 2019.  
Reproductionoviparous 
TypesHolotype: MZUC 8578 (Concepción, Chile), male, fide Murphy et al. 2019, was: DB 6312061, i.e. private collection of Donoso-Barros, collected on 12 December 1963 by R. Donoso-Barros and A. Garrido. Paratype: female, probably also in MZUC. 
DiagnosisDiagnosis A Bachia with (1) no prefrontal scales; (2) two supraoculars; (3) the frontonasal is hexagonal; (4) four digits are present on the manus and two on the pes (5) a band of imbricate hexagonal scales on the dorsum; (6) scales around mid-body 25; (7) 39–42 transverse rows of dorsals; (8) the fifth upper labial contacts primary temporal; (9) the gulars are in five rows; (10) the second pair of chin shield are in medial contact; (11) a sub-rectangular interparietal present (from Murphy et al. 2019). 
CommentSynonymy partly that of Dixon (1973) who synonymized B. marcelae with B. heteropa. However, more recent authors recognized marcelae as valid. 
EtymologyNamed after Marcela, one of Donoso-Barros’ daughters (althought not given by Donoso-Barros 1964). 
References
  • Bauer, A.M. and R. Günther 1994. An annotated type catalogue of the teiid and microteiid lizards in the Zoological Museum, Berlin (Reptilia: Squamata: Teiidae and Gymnophthalmidae). Mitt. Zool. Mus. Berlin 70:267-280 - get paper here
  • Bentz, Ehren J.; Mel José Rivera Rodríguez, Rebecca R. John, Robert W. Henderson, and Robert Powell 2011. Population Densities, Activity, Microhabitats, and Thermal Biology of a Unique Crevice- and Litter-dwelling Assemblage of Reptiles on Union Island, St. Vincent and the Grenadines. Herp. Cons. Biol. 6 (1): 40-50 - get paper here
  • Boos, H.E.A. 1984. The terrestrial reptiles of Monos island. Living World - get paper here
  • Daan, S. & Hillenius,D. 1966. Catalogue of the type specimens of amphibians and reptiles in the Zoological Museum, Amsterdam. Beaufortia 13: 117-144
  • Daudin, Jacques; de Silva, Mark. 2007. An annotated checklist of the amphibians and terrestrial reptiles of the Grenadines with notes on their local natural history and conservation. Applied Herpetology 4 (2):163-175 - get paper here
  • Disi, Ahmad; Zuhair S. Amr, Nashat Hamidan 2014. Diversity, Threats, and Conservation of the Terrestrial and Freshwater Herpetofauna of Jordan. Russ. J. Herpetol. 21 (3): 221-233 - get paper here
  • Dixon, J.R. 1973. A systematic review of the teiid lizards, genus Bachia, with remarks on Heterodactylus and Anotosaura. Misc. Publ. Univ. Kans. Mus. Nat. Hist. 57: 1-47 - get paper here
  • Donoso-Barros, R. 1968. The lizards of Venezuela (checklist and key). Carib. J. Sci. 8 (3-4): 105-122
  • Donoso-Barros, R. and A. Garrido R. 1964. Nuevo Teiidae de Venezuela, Bachia marcelae, nov. sp. Publ. Ocas. Mus. Cienc. Nat., Caracas (Zool.) 8: 1-7
  • Esqueda, Luis Felipe, Enrique La Marca, Marco Natera and Pietro Battiston 2001. Noteworthy reptilian state records and a lizard species new to the herpetofauna of Venezuela. Herpetological Review 32 (3):198-200 - get paper here
  • Gorzula, Stefan & Senaris, J. Celsa 1999. In: Contribution to the herpetofauna of the Venezuelan Guayana. I: a data base. Scientia Guaianae, Caracas, No. 8 [1998], 269+ pp.; ISBN 980-6020-48-0
  • John, Rebecca R., Ehren J. Bentz, Mel Jose Rivera Rodriguez, Aaron M. Bauer and Robert Powell. 2012. Bachia heteropa. Catalogue of American Amphibians and Reptiles (894): 1-9 - get paper here
  • Kramer, Eugen 1979. Typenkatalog der Echsen im Naturhistorischen Museum Basel (BM), Stand 1978. [type catalogue] Revue Suisse de Zoologie 86 (1): 159-166 - get paper here
  • Markezich, A.L. 2002. New distribution records of reptiles from Western Venezuela. Herpetological Review 33 (1): 69-74 - get paper here
  • Mijares-Urrutia, A. & Arends R., Alexis 1999. Additional New Regional and Local Records of Amphibians and Reptiles from the State of Falcón, Venezuela. Herpetological Review 30 (2): 115 - get paper here
  • Murphy, John C.; Daniele Salvi, Joana L. Santos, Alvin L. Braswell, Stevland P. Charles, Amaél Borzée & Michael J. Jowers 2019. The reduced limbed lizards of the genus Bachia (Reptilia, Squamata, Gymnophthalmidae); biogeography, cryptic diversity, and morphological convergence in the eastern Caribbean. Organisms Diversity & Evolution - get paper here
  • Powell, Robert and Robert W. Henderson 2017. Notes on the diets of some amphibians and reptiles of the St. Vincent and Grenada Banks (Lesser Antilles). IRCF Reptiles & Amphibians 24 (2): 135–138 - get paper here
  • Ribeiro-Júnior, Marco A.; Marcélia B. da Silva, and Jucivaldo D. Lima 2016. A New Species of Bachia Gray 1845 (Squamata: Gymnophthalmidae) from the Eastern Guiana Shield. Herpetologica 72 (2): 148-156. - get paper here
  • Rivas Fuenmayor, G.; Ugueto, G.; Rivero, R. & Miralles, A. 2005. The Herpetofauna of Isla de Margarita, Venezuela: New Records and Comments. Carib. J. Sci. 41 (2): 346-351
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Señaris, J. Celsa; María Matilde Aristeguieta Padrón, Haidy Rojas Gil y Fernando J. M. Rojas-Runjaic 2018. Guía ilustrada de los anfibios y reptiles del valle de Caracas, Venezuela. Ediciones IVIC, Instituto Venezolano de Investigaciones Científicas (IVIC). Caracas, Venezuela. 348 pp.
  • Vanzolini,P.E. 1961. Bachia: Especies Brasileiras e conceito generico (Sauria: Teiidae). Papéis Avulsos Zoologia (São Paulo) 14: 193-209 - get paper here
 
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