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Bachia remota RIBEIRO-JÚNIOR, DA SILVA & LIMA, 2016

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Higher TaxaGymnophthalmidae (Cercosaurinae, Bachiinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Common Names 
SynonymBachia remota RIBEIRO-JÚNIOR, DA SILVA & LIMA 2016
Bachia gr. heteropa — LIMA 2008: 42 
DistributionBrazil (Amapá)

Type locality: Parque Nacional Montanhas do Tumucumaque (2°11’36’’N, 54°35’15’’W; datum 5 WGS84), Laranjal do Jari municipality, State of Amapá, Brazil. The park is located on the Brazilian border with French Guiana and Suriname  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: IEPA 777 (Figs. 1–3), an adult collected on 20 January 2005 by J. Lima. Field number TQ 372. 
DiagnosisDiagnosis: Bachia remota is a member of the heteropa group with hexagonal, smooth, imbricate dorsal, and rectangular, juxtaposed ventral and lateral scales. The new species is distinguished from all other species in the heteropa group by the following combinations of characters: (1) four clawed digits on all limbs; (2) prefrontals absent; (3) interparietal absent; (4) two supraciliary scales; (5) large size (86.8 mm SVL); and, (6) background coloration homogeneous brown on all surfaces of body and tail.

Comparisons: Bachia remota differs from all species of the B. bresslaui group (B. bresslaui; B. cacerensis; B. didactyla; B. geralista; B. micromela; B. oxyrhina; B. panoplia; B. psamophila; B. pyburni; and B. scolecoides) by having hexagonal, smooth, imbricate dorsal, and rectangular, juxtaposed lateral scales (vs. keeled dorsal, and hexagonal lateral scales in all other species). Differs from the B. dorbignyi group (B. dorbignyi); B. barbouri; B. bicolor; B. huallagana; B. intermedia; B. peruana; B. scaea; B. talpa; and B. trisanale by having rectangular, juxtaposed lateral scales, four clawed digits on each limb, and supraoculars present (vs. hexagonal lateral scales, three digits on each limb, and supraoculars absent in all other species). Differs from B. flavescens by having hexagonal, imbricate dorsal (vs. quadrangular dorsal scales).
Considering the species of the Bachia heteropa group (Table 1), B. remota can be distinguished from B. heteropa and B. pallidiceps by the absence of interparietal scale, and having 56 dorsal scales (vs. presence of interparietal, and 38–49 dorsal scales in B. heteropa, 43–48 in B. pallidiceps), and from B. guianensis and B. pallidiceps by the absence of prefrontal scales. Bachia remota can also be distinguished from B. guianensis by having two supraciliary scales and a homogeneous brown body (vs. three supraciliaries, and dorsal surface of body and tail with four longitudinal series of white spots). The new species can be distinguished from other species occurring in Amazonia (Table 2 in Ribeiro-Júnior et a. 2016). 
CommentDistribution: see map in Ribeiro-Júnior et al. 2016: 153 (Fig. 4).

Abundance: only known from the type specimen (Meiri et al. 2017). 
EtymologyThe specific epithet is derived from the Latin adjective remotus and refers to the geographical remoteness and isolation of the region (Montanhas do Tumucumaque National Park) where the specimen was collected. 
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • RIBEIRO-JÚNIOR, MARCO A. & SILVANA AMARAL 2017. Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. IV. Alopoglossidae, Gymnophthalmidae. Zootaxa 4269 (2): 151-196 - get paper here
  • Ribeiro-Júnior, Marco A.; Marcélia B. da Silva, and Jucivaldo D. Lima 2016. A New Species of Bachia Gray 1845 (Squamata: Gymnophthalmidae) from the Eastern Guiana Shield. Herpetologica 72 (2): 148-156. - get paper here
  • Ribeiro-Júnior, Marco A. & Silvana Amaral 2016. Diversity, distribution, and conservation of lizards (Reptilia: Squamata) in the Brazilian Amazonia. Neotropical Biodiversity, 2:1, 195-421 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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