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Blaesodactylus microtuberculatus JONO, BAUER, BRENNAN & MORI, 2015

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymBlaesodactylus microtuberculatus JONO, BAUER, BRENNAN & MORI 2015
Blaesodactylus microtuberculatus — INEICH et al. 2016 
DistributionN Madagascar (Antsiranana)

Type locality: Ankarana National Park, Antsiranana Province, northern Madagascar (12°57’S, 49°07’E; Fig. 3),  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: KUZ 069431 (field number AFP2012-002), adult male (Fig. 2A); collected by T. Jono and A. Mori on 4 December 2012. Paratypes. None. 
DiagnosisDiagnosis: A large sized (SVL 117.4 mm), robust-bodied Blaesodactylus with tail slightly longer than SVL (Fig. 2A). Snout elongate, gular granules small and homogenous, internarial region convex (Fig. 4A). 162 scale rows around midbody, 15–17 longitudinal rows of relatively small tubercles on dorsum. Fourth digit of pes with 21 undivided subdigital lamellae. No tubercles on tail dorsum; midventral subcaudal scales transversely enlarged, about half width of tail. Body dorsum grayish, mid-dorsal row of five blotches, original tail weakly banded, venter pale, only gular region with very faint mottling (Fig. 4A).
Among its congeners Blaesodactylus microtuberculatus sp. nov. may be differentiated from B. boivini by its smaller dorsal tubercles (Fig. 5; ~3 times size of adjacent scales versus ~10 times), absence of caudal tubercles (Fig. 5; versus caudal tubercles ~4 times size of adjacent scales and present on more than half of the tail), uniform pale venter except for gular region (versus mottled with areas of dark pigmentation), and lower number of dorsal tubercle rows (15–17 versus 18–21). It differs from B. sakalava in its smaller dorsal tubercles (Fig. 5; ~3 times size of adjacent scales versus ~6 times, with tubercles finely keel), absence of caudal tubercles (Fig. 5; versus keeled tubercles ~6 times size of adjacent scales and present on more than half of the tail), wider midventral subcaudal scales (Fig. 5; about half width of tail versus less than one-third width of tail), and mottled gular region (Fig. 4; versus pale). It is distinguished from B. ambonihazo by its smaller dorsal tubercles (Fig. 5; ~3 times size of adjacent scales versus ~6 times), absence of caudal tubercles (Fig. 5; versus keeled tubercles ~3 times size of adjacent scales present on proximal ~40% of the tail), wider midventral subcaudal scales (Fig. 5; about half width of tail versus less than one-third width of tail), mottled gular region (Fig. 4; versus pale), lower number of dorsal tubercle rows (15–17 versus 17–21). It is distinguished from B. antongilensis by its lower number of dorsal tubercles (15–17 versus up to 24), and mottled gular region (Fig. 4; versus pale). [JONO et al. 2015]. 
CommentAbundance: only known from the type locality (Meiri et al. 2017). 
EtymologyThe species epithet “microtuberculatus” refers to the presence of small tubercles, compared to all other congeners, on their trunk region. 
References
  • Ineich, I.; FRANK GLAW, MIGUEL VENCES 2016. A new species of Blaesodactylus (Squamata: Gekkonidae) from Tsingy limestone outcrops in Namoroka National Park, north-western Madagascar. Zootaxa 4109 (5): 523–541 - get paper here
  • JONO, TEPPEI; AARON M. BAUER, IAN BRENNAN & AKIRA MORI 2015. New species of Blaesodactylus (Squamata: Gekkonidae) from Tsingy karstic outcrops in Ankarana National Park, northern Madagascar. Zootaxa 3980 (3): 406–416 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Rösler, Herbert 2017. Gecko-Chorologie (Squamata: Gekkota). Gekkota (4): 1-160
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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