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Bothriechis klebbai ARTEAGA, PYRON, BATISTA, VIEIRA, MENESES-PELAYO, SMITH, BARRIO-AMORÓS, KOCH, AGNE, VALENCIA, BUSTAMANTE & HARRIS, 2024

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Klebba’s Eyelash-Pitviper
G: Klebbas Lanzenotter
S: Víbora de pestañas de Klebba 
SynonymBothriechis klebbai ARTEAGA, PYRON, BATISTA, VIEIRA, MENESES-PELAYO, SMITH, BARRIO-AMORÓS, KOCH, AGNE, VALENCIA, BUSTAMANTE, HARRIS 2024: 41 
DistributionColombia (Cordillera Oriental)

Type locality: Vereda Virolín, municipio de Charalá, Santander department, Colombia (6.10720, -73.19781; 1819 m elevation.  
Reproduction 
TypesHolotype: ICN 2786 (Fig. 19), adult male collected by Pedro M. Ruiz-Carranza on September 21, 1979.
Paratypes: See Suppl. material 1 in Arteaga et al. 2024. 
DiagnosisDiagnosis: Bothriechis klebbai sp. nov. is diagnosed based on the following combination of characters: (1) supraciliary scales low and granular or two raised, but not sharp; (2) anterior dorsal head scales smooth; (3) gular scales similar in size to chinshields; (4) 7–11 interoculolabials; (5) 2–4 canthals which may be flat or slightly raised forming a ridge along the canthus; (6) loreal in contact with preocular in ~1/3 of specimens; (7) yellow morph absent; (8) dorsal bands black; (9) opposing kidney shaped dorsal marks absent; (10) black speckles on dorsal scales present; (11) black speckling on ventral surfaces present, prominent; (12) ventral surfaces never entirely white; (13) iris rich dark reddish brown; (14) 21–23 dorsal scale rows at mid-body; (15) 144–151 ventrals in males, 141–151 in females; (16) maximum total length in males 671 mm, in females 874 mm. (Arteaga et al. 2024: 41) 
CommentVenomous!

Synonymy: Reyes-Velasco 2024 argued that B. torvus, B. khwargi, B. klebbai, B. rasikusumorum, B. rahimi, B. nitidus, B. torvus, and B. hussaini should be synonymized with B. schlegelii, given their genetic and morphological similarity. Reyes-Velasco 2024 thus argued that only B. nigroadspersus, B. schlegelii, and B. supraciliaris should be maintained. We agree that the former 5 species are not very much differentiated from B. schlegelii and should be considered preliminary species. We have not adopted Reyes-Velasco’s revision yet as there are reportedly other studies in the process of publication, hence the taxonomy of Bothriechis is in flux and more changes are likely to come soon. 
References
  • Arteaga A, Pyron RA, Batista A, Vieira J, Meneses Pelayo E, Smith EN, Barrio Amorós CL, Koch C, Agne S, Valencia JH, Bustamante L, Harris KJ 2024. Systematic revision of the Eyelash Palm-Pitviper Bothriechis schlegelii (Serpentes, Viperidae), with the description of five new species and revalidation of three. Evolutionary Systematics 8(1): 15-64 - get paper here
  • Reyes-Velasco, J. 2024. A revision of recent taxonomic changes to the eyelash palm pitviper, Bothriechis schlegelii (Serpentes, Viperidae). Herpetozoa, 37: 305-318 - get paper here
 
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