Bothrops chloromelas (BOULENGER, 1912)

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Higher Taxa	Viperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names
Synonym	Lachesis chloromelas BOULENGER 1912: 423 Bothrops chrysomelas AMARAL 1926 (in error for chloromelas) Bothrops chloromelas— WÜSTER et al. 2002 Bothriopsis chloromelas — CAMPBELL & LAMAR 2004 Bothriopsis chloromelas — HARVEY et al. 2005 Bothriopsis chloromelas — FENWICK et al. 2009 Bothrops chloromelas — CARRASCO et al. 2012 Bothriopsis chloromelas — WALLACH et al. 2014: 109
Distribution	Peru (central Andes) Type locality: “Huancabamba, E. Peru, above 3000 feet” [elevation]
Reproduction	ovovivparous
Types	Syntypes: BMNH 1946.1.17.66 (formerly BMNH 1911.12.13.63) and BMNH 1946.1.19.51-52.
Diagnosis	Diagnosis: Bothriopsis chloromelas is distinguished from all congeners and from species of Bothrocophias and Bothrops by the following combination of characters: (1) lacunolabial present; (2) dorsal keels nontuberculate; (3) infralabials yellow with small black spots; (4) canthoros- trals absent; (5) anteriormost 5–25 subcaudals entire; dis- tal subcaudals divided; (6) ventrals 183–204 and subcau- dals 49–64; (7) loreal subtriangular; (8) canthals narrow; (9) postocular stripe black and 2–3 temporals high; (11) head black with yellow lines forming circular patterns or head yellow with heavy black stippling and marking; supralabials green, heavily speckled in black; dorsum banded and lichenose (Harvey 2005: 29).
Comment	Lachesis chloromelas BOULENGER 1912 has been revalidated by HARVEY 2004, a notion that also CAMPBELL & LAMAR (2004) expressed without detailed justification. Habitat: fully arboreal (Harrington et al. 2018).
Etymology	Named after Greek chloros (χλωρός), yellow; greenish yellow; green; pale, pallid. + Greek mélas (μέλας), black [“…Yellowish green above, speckled with black, with large black irregular spots...”]. (from Esteban Lavilla, pers. comm., May 2024)
References	Amaral,A. do 1926. Annals of the Carnegie Museum 16: 320, 322 Boulenger, George A. 1912. Descriptions of new reptiles from the Andes of South America, preserved in the British Museum. Ann. Mag. nat. Hist. (8) 10: 420-424 - get paper here Campbell, Jonathan A. and William W. Lamar 2004. The Venomous Reptiles of the Western Hemisphere, 2 vols. Comstock (Cornell University Press), Ithaca, NY, 962 pp. [review in Science 305: 182] Carrasco, P.A., C.I. Mattoni, G.C. Leynaud, and G.J. Scrocchi. 2012. Morphology, phylogeny and taxonomy of South American bothropoid pitvipers (Serpentes, Viperidae). Zoologica Scripta 41: 109-124 - get paper here Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here Harvey, M.B.; et al. 2005. Revision of the venomous snakes of Bolivia. II. The pitvipers (Serpentes: Viperidae). Annals of the Carnegie Museum 74 (1): 1-38. - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
External links	IUCN Red List National Center for Biotechnology Information http://biology.bangor.ac.uk/~bss166/Updates/Crotalinae2005.htm Google images

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