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Brachymeles miriamae (HEYER, 1972)

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Higher TaxaScincidae, Scincinae, Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Miriam's Skink
G: Miriams Skink 
SynonymDavewakeum miriamae HEYER 1972
Davewakeum miriamae — GREER 2002
Brachymeles miriamae — SILER et al. 2011
Davewakeum miriamae — CHAN-ARD et al. 2015: 136
Brachymeles miriamae — COTA et al. 2022 
DistributionThailand (Nakhon Ratchasima Province)

Type locality: Khao Saton, 300 m above the Kasetsart University Forestry station, 60 km S of Nakhon Ratchasima on Highway 304, Amphoe Pak Thong Chai, Changwat Nakhon Ratchasima, Thailand.  
Reproductionviviparous (phylogenetic imputation, fide Zimin et al. 2022) 
TypesHolotype: FMNH 182550, but THNHM 20240 according to Cota et al. 2022. 
DiagnosisDiagnosis (genus and species): The only scincine genera in which some or all of the species are limbless are: Barkudia, Brachymeles, Cryptoposcincus, Davewakeum,Grandidierina, Malacontias, Melanoseps, Nessia, Ophiomorus, Paracontias, Pseudoacontias, Scelotes, Scolecoseps, Sepsophis, Typhlacontias, Voeltzkowia. Barkudia and Sepsophis differ from Davewakeum in having an external ear. The remainder of the genera either lack an external ear or the character is variable among species within a genus. Ophiomorus has a transparent disk in the lower eyelid. Davewakeum has a thickened, opaque lower eyelid. Brachymeles, Melanoseps, Scelotes, Scolecoseps, and Typhlacontias have supranasal scales:Davewakeum lacks supranasal scales. Cryptoposcincus, Grandidierina, Malacontias, Nessia, Paracontias, Pseudoacontias, and Voeltzkowia lack prefrontal scales, Davewakeum has prefrontal scales [from HEYER 1972].


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CommentLimb morphology: Limbless.

Type Species: Davewakeum miriamae HEYER 1972 is the type species of the genus Davewakeum HEYER 1972, which nests within Brachymeles, at least when B. apus is included (e.g. Zheng & Wiens 2016).

Habitat: semifossorial and typically found in dry, rotting material inside or underneath decaying logs or in loose soil, forest floor detritus, and leaf litter of lowland forest. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Cota, M., Makchai, S., & Pongcharoen, C. 2022. Updated Catalogue of the Amphibian and Reptile Type Specimens of the Natural History Museum, National Science Museum, Thailand. Thai Specimens 21: 311––5407
  • Greer, A.E. 2002. The loss of the external ear opening in scincid lizards. Journal of Herpetology 36 (4): 544-555 - get paper here
  • Heyer, W.R. 1972. A new limbless skink (Reptilia: Scincidae) from Thailand with comments on the generic status of the limbless skinks of southeast Asia. Fieldiana: Zoology 58 (10): 109-129 - get paper here
  • Siler, Cameron D.; Arvin C. Diesmos, Angel C. Alcala and Rafe M. Brown 2011. Phylogeny of Philippine slender skinks (Scincidae: Brachymeles) reveals underestimated species diversity, complex biogeographical relationships, and cryptic patterns of lineage diversification. Molecular Phylogenetics and Evolution 59 (1): 53-65 - get paper here
  • Staub, Nancy L. and Rachel Lockridge Mueller 2017. David Burton Wake. Copeia 105 (2): 415-426 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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