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Broadleysaurus major (DUMÉRIL, 1851)

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Higher TaxaGerrhosauridae (Gerrhosaurinae), Scincoidea, Sauria, Squamata (lizards) 
Subspecies 
Common NamesE: Rough-scaled Plated Lizard
G: Braune Schildechse, Sudan-Schildechse 
SynonymGerrhosaurus major major DUMÉRIL 1851
Gerrhosaurus major DUMÉRIL 1851
Gerrhosaurus zanzibaricus PFEFFER 1889
Gerrhosaurus bottegoi DEL PRATO 1895
Gerrhosaurus major var. zechi TORNIER 1901
Gerrhosaurus bergi WERNER 1906
Gerrhosaurus grandis BOULENGER 1908
Gerrhosaurus major grandis — HEWITT 1910
Gerrhosaurus zechi SCHMIDT 1919: 519
Gerrhosaurus major zechi — LOVERIDGE 1936: 64
Gerrhosaurus major zechi — MERTENS 1938
Gerrhosaurus major bottegoi — PARKER 1942: 77
Gerrhosaurus major bottegoi - BROADLEY 1987
Gerrhosaurus major major - BROADLEY 1987
Gerrhosaurus major major — AUERBACH 1987: 118
Gerrhosaurus major major — BOYCOTT 1992
Gerrhosaurus major major — BROADLEY & HOWELL 1991: 18
Gerrhosaurus major — LARGEN & SPAWLS 2006: 43
Gerrhosaurus major zechi — CHIRIO & LEBRETON 2007
Gerrhosaurus major bottegoi — BARTS 2008
Gerrhosaurus major zechi — ULLENBRUCH et al. 2010
Gerrhosaurus major bottegoi — TRAPE, CHIRIO & TRAPE 2012
Broadleysaurus major — BATES et al. 2013
Broadleysaurus major bottegoi — NCBI TAXONOMY 2016 
DistributionE Kenya, Tanzania, Mozambique, Malawi, N Zambia,
Ethiopia, Eritrea, Togo, N Democratic Republic of the Congo (Zaire), Republic of South Africa, Swaziland, Botswana,
Zimbabwe, S Sudan, Central African Republic, Somalia,
Cameroon, Nigeria, Benin, Ghana

bottegoi: Central African Republic, Somalia, Djibouti

zechi: Benin, Cameroon

Type locality: Zanzibar Island.
Type locality: Zululand [grandis] Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesSyntype: MNHN
Holotype: ZMB 18814 [bergi]
Syntypes: ZMB 16608 [zechi] 
CommentSynonymy: BATES et al. (2013) synonymized Gerrhosaurus (major) bottegoi with G. (now Broadleysaurus) major.

NCBI taxonomy: 219621 (bottegoi)

Diagnosis (genus): The monophyly of Broadleysaurus is established on the basis of a suite of nuclear and mitochondrial genetic characters (see above). This genus of terrestrial gerrhosaurids is differentiated from members of the genus Gerrhosaurus and the new genus described below for ‘G. validus’ by the following morphological characters: Body stout, cyclotetragonal to feebly depressed dorso-ventrally, and well armoured; readily distinguished from the above-named genera by its much lower (31–38 versus 49–67) numbers of transverse dorsal scale rows; also distinguished by having 9–10 longitudinal rows of ventrals (8 in Gerrhosaurus, but 10 in G. typicus; 12–20 in the new genus described in Bates et al. 2013 for ‘G. validus’) (Loveridge 1942; FitzSimons 1943, 1953; De Witte 1953; Laurent 1954, 1964; Broadley 1966; De Waal 1978; Jacobsen 1989) [BATES et al. 2013].

Type species: Gerrhosaurus major DUMÉRIL 1851 is the type species of the genus Broadleysaurus BATES & TOLLEY 2013. 
EtymologyThe genus has been named in honour of Dr Donald George Broadley, Curator Emeritus at the Natural History Museum, Bulawayo (Zimbabwe) for his numerous contributions to African herpetology (e.g. see Branch 1997). He was recently ranked as the third most productive living author of reptile names (Uetz 2010). It is also fitting to honour Dr Broadley with this name as his review (1987) of ‘Gerrhosaurus major’ is the most recent and authoritative. The name Broadleysaurus is masculine, derived from Broadley + the Latinised word saurus meaning lizard. 
References
  • Auerbach, R.D. 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Botswana, 295 pp.
  • Barnes, Keith 2016. Animals of Kruger National Park. Princeton University Press, ca. 180 pp. - get paper here
  • Barts, M. 2008. Schildechsen: Gattungen und Arten im Überblick. Reptilia (Münster) 13 (69): 18-25 - get paper here
  • BATES, MICHAEL F.; KRYSTAL A. TOLLEY, SHELLEY EDWARDS, ZOË DAVIDS, JESSICA M. DA SILVA, & WILLIAM R. BRANCH 2013. A molecular phylogeny of the African plated lizards, genus Gerrhosaurus Wiegmann, 1828 (Squamata: Gerrhosauridae), with the description of two new genera. Zootaxa 3750 (5): 465–493 - get paper here
  • Bonetti, Mathilde 2002. 100 Sauri. Mondadori (Milano), 192 pp.
  • Boulenger, G. A. 1908. On a collection of fresh-water fishes, batrachians, and reptiles from Natal and Zululand, with description of new species. Ann Natal Gov. Mus. (Pietermaritzburg) 1(3): 219-235
  • Boycott, R.C. 1992. An Annotated Checklist of the Amphibians and Reptiles of Swaziland. The Conservation Trust of Swaziland - get paper here
  • Branch, William R. 1993. A Photographic Guide to Snakes and Other Reptiles of Southern Africa. Cape Town: Struik Publishers, 144 S.
  • Broadley, D. G. 1987. A review of Geographical variation in Gerrhosaurus major Dumeril (Sauria: Cordylidae). The Herpetological Journal 1: 194-198
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1962. On some reptile collections from the North-Western and North-Eastern Districts of Southern Rhodesia 1958-1961, with descriptions of four new lizards. Occ. Pap. Nat. Mus. South. Rhodesia 26 (B): 787-843
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Chirio, Laurent and Ivan Ineich 2006. Biogeography of the reptiles of the Central African Republic. African Journal of Herpetology 55(1):23-59. - get paper here
  • Dassow, S. 2008. Ein kleiner Bericht über die Sudan-Schildechse (Gerrhosaurus major). Reptilia (Münster) 13 (69): 32-36 - get paper here
  • Dathe, Falk 1987. AT Terrarientierlexikon, Gerrhosaurus major Duméril, 1851, Braune oder Sudan-Schildechse (Familie: Cordylidae, Gürtel- und Schildechsen). Aquarien und Terrarien (Leipzig), 34 (12): 423.
  • del Prato, Alberto 1895. Vertebrati Eritri aggiunta al catalogo della collezione Eritrea Bottego. Atti. Soc. Ital. Sci. Nat. (Milano) 35: 17-26
  • Duméril, A.M.C. & A. H. A. Duméril 1851. Catalogue méthodique de la collection des reptiles du Muséum d'Histoire Naturelle de Paris. Gide et Baudry/Roret, Paris, 224 pp.
  • Ineich, I. 1999. Reptiles & Amphibiens de la République de Djibouti. BNDB, UICN, PNUD, Djibouti & MNHN, Paris, 60 pp. - get paper here
  • Jacobsen, Niels H.G.; Errol W. Pietersen & Darren W. Pietersen 2010. A preliminary herpetological survey of the Vilanculos Coastal Wildlife Sanctuary on the San Sebastian Peninsula, Vilankulo, Mozambique. Herpetology Notes 3: 181-193 - get paper here
  • John, Wolfgang 1980. Beobachtungen im Biotop der Brauen [sic!] oder Sudan-Schildechse (Gerrhosaurus major) A. Duméril 1851. Herpetofauna 2 (5): 31-32. - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988]
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  • Leaché, Adam D.; Mark-Oliver Rödel, Charles W. Linkem, Raul E. Diaz, Annika<br />Hillers, and Matthew K. Fujita 2006. Biodiversity in a forest island: reptiles and amphibians of the Togo Hills, Kyabobo National Park, Ghana. Amphibian & Reptile Conservation 4 (1): 22-45 - get paper here
  • Loveridge, A. 1923. Notes on East African lizards collected 1920-1923 with the description of two new races of Agama lionotus Blgr. Proc. Zool. Soc. London 1923: 935-969 - get paper here
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Loveridge,A. 1920. Notes on East African lizards collected 1915-1919, with description of a new genus and species of skink and new subspecies of gecko. Proc. Zool. Soc. London 1920: 131-167 - get paper here
  • Meek, Seth Eugene;Cory, Charles B. 1910. Batrachians and reptiles from British East Africa. Publication of the Field Museum of Natural history 7 (11): 403-414 - get paper here
  • Mertens, R. 1938. Über eine herpetologische Sammlung aus dem Gebiete des Njarasa-Grabens Ost-Afrikas. Veröffentlichungen aus dem Deutschen Kolonial- und Übersee-Museum in Bremen 2 (1): 1-9 [no page numbers]
  • Parker, H. W. 1942. The lizards of British Somaliland. Bull. Mus. Comp. Zool. Harvard 91: 1—101 - get paper here
  • Pfeffer, G. 1889. Übersicht der von Herrn Dr. Franz Stuhlmann in Ägypten, auf Sanzibar und dem gegenüberliegenden Festlande gesammelten Reptilien, Amphibien, Fische, Molluskten und Krebse. Jb. Hamb. Wiss. Anst. 6 (2): 1-36
  • Pfeffer, G. 1893. Ostafrikanische Reptilien und Amphibien, gesammelt von Herrn Dr. F. Stuhlmann im Jahre 1888 und 1889. Jb. Hamb. Wiss. Anst. 10: 71-105.
  • Rese, Rolf 1981. Schildechsen (Gerrhosaurus) - Haltung und Zucht. Sauria 3 (3): 29-32 - get paper here
  • Rese, Rolf 1986. Gerrhosaurus major Duméril. Sauria 8 (4) Suppl.: 61-62. - get paper here
  • Schmidt, Karl Patterson 1919. Contributions to the Herpetology of the Belgian Congo based on the Collection of the American Congo Expedition, 1909-1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bull. Amer. Mus. Nat. Hist. 39 (2): 385-624 - get paper here
  • Segniagbeto, Gabriel Hoinsoude; Jean-François Trape, Komlan M. Afiademanyo, Mark-Oliver Rödel, Annemarie Ohler, Alain Dubois, Patrick David, Danny Meirte, Isabelle Adolé Glitho, Fabio Petrozzi, and Luca Luiselli 2015. Checklist of the lizards of Togo (West Africa), with comments on systematics, distribution, ecology, and conservation. Zoosystema 37 (2): 381-402 - get paper here
  • Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83 - get paper here
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Trape, J.F.; Trape, S. & Chirio, L. 2012. Lézards, crocodiles et tortues d'Afrique occidentale et du Sahara. IRD Orstom, 503 pp. - get paper here
  • Ullenbruch, K.; Grell, O.; Böhme, W. 2010. Reptiles from southern Benin, West Africa, with the description of a new Hemidactylus (Gekkonidae), and a country-wide checklist. Bonn Zool. Bull. 57 (1): 31-54 - get paper here
  • Werner, Franz Josef Maria 1906. Neue Reptilien aus Deutsch-Ostafrika. Zool. Anz. 30: 53-55. - get paper here
 
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