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Bronchocela cristatella (KUHL, 1820)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Green crested lizard
G: Borneo-Langschwanzagame, Borneo-Schönechse, Blutsaugeragame 
SynonymAgama cristatella KUHL 1820: 108
Agama gutturosa MERREM 1820
Bronchocela cristatella — KAUP 1827: 619
Bronchocela gutturosa — KAUP 1827: 619
Calotes gutturosa — CUVIER 1829: plate 7
Agama moluccana LESSON 1830: 34
Calotes cristatellus — WIEGMANN in MEYEN 1834: 219
Bronchocela cristatella — DUMÉRIL & BIBRON 1837: 395
Pseudocalotes archiducissae FITZINGER 1861 (nom. nud. fide SMITH 1935)
Calotes (Bronchocele) philippinus PETERS 1867: 16 (part.)
Bronchocela intermedia PETERS & DORIA 1878: 375
Calotes cristatellus — BOULENGER 1885: 316
Calotes cristatellus — FLOWER 1899
Calotes octospinosus BAUMANN 1913
Calotes cristatellus — DE ROOIJ 1915: 121
Calotes cristatellus — TAYLOR 1922: 139
Calotes cristatellus — SMITH 1935: 184
Calotes cristatellus moluccanus — BRONGERSMA 1948: 489
Calotes cristatellus moluccanus — TANNER 1950
Calotes cristatellus — TAYLOR 1963: 886
Calotes cristatellus — HENDRICKSON 1966: 64
Bronchocela cristatella — MANTHEY 1983: 25
Bronchocela petersdoriai BÖHME & BISCHOFF 1984 (nom. nov. pro B. intermedia)
Calotes cristatellus — BONG HEANG 1987
Calotes cristatellus — BROWN et al. 1996
Bronchocela cristatella — MANTHEY & GROSSMANN 1997: 160
Bronchocela cristatella — COX et al. 1998: 98
Bronchocela cristatella — GAULKE 1999
Bronchocela christatella — CHAN-ARD et al. 1999 (in error)
Bronchocela cf. cristatella — GAULKE 2001
Bronchocela cristatella — BARTS & WILMS 2003
Bronchocela cristatella — NUR-AMALINA et al. 2017
Bronchoecella cristatella — BADLI-SHAM et al. 2019 (in error)
Bronchocela cf. cristatella — GLÄSSER-TROBISCH & TROBISCH 2021 
DistributionMalaysia (Pulau Pangkor, Pulau Tioman, Pulau Langkawi, Pulau Singa Besar, Johor: Pulau Sibu, Borneo),
S Myanmar (= Burma) (Tenasserim), S Thailand, W Malaysia, Singapore, India (Andaman and Nicobar Islands), Philippines (Palawan, Calamian Islands, Panay, Luzon, Mindanao, Sulu, Cebu, Bohol, Leyte, Negros, Samar, Mindoro, Masbate), Indonesia (Halmahera, Sumatra, Morotai, Natuna islands etc.), Papua New Guinea

Type locality: not given by KUHL.

Type locality: “Kandari nel S.E. di Selebes” [= Kendari, Sulawesi, Indonesia] [Bronchocela intermedia PETERS & DORIA 1878].  
Reproductionoviparous 
TypesLectotype (iconotype): Seba (1734: plate 89 figure 1 (designated by Figueroa 2021); this is a lectotype designation for both A. cristatella and Agama gutturosa.
Holotype: lost fide Brygoo 1988 but Das & Gemel 2000 suggested that KUHL based his name on MNHN 6884 (three syntypes). Other specimens: RMNH, ZFMK, ZMH, MZB, ZSM
Holotype: NMW 21002: 1-2 [archiducissae] 
DiagnosisDiagnosis (genus): Bronchocela species are slender, medium sized (SVL about 80–140 mm) with long tails TL/SVL 3,2–3,9, laterally compressed body and relatively long legs. The genus can be separated from all other draconinae genera by a fold of skin that extends from the angle of the jaw to the insertion of the upper arm; homogeny lateral scales rhombic and small, point backwards and downwards, ventral scales greater; tympanum naked; nuchal crest present, lower dorsal crest sometimes reduced to a serrated ridge; spines on the head absent (U. Manthey, pers. comm., 3 Aug 2021). 
CommentDistribution: According to SMITH 1935 not on New Guinea. See map in Amarasinghe et al. 2022: 411 (Fig. 1)

Type species: Agama gutturosa MERREM 1820 is the type species of the genus Bronchocela KAUP 1827: 619, designated by Fitzinger 1843 (not A. cristatella, as sometime stated in the literature, see Figueroa 2021 for a discussion).

Synonymy: Boulenger (1880) synonymized B. burmana with Calotes cristatellus but Zug et al. 2017 revalidated Bronchocela burmana again. Bronchocela intermedia PETERS & DORIA 1878 is a junior homonym of B. intermedia BERTHOLD 1842.

The Bronchocela cristatella mentoned in ter Borg (2005) is probably B. jubata (ter Borg 2007).

Comparisons: for a comparison of diagnostic characters across the species of Bronchocela see Amarasinghe et al. 2022: 73 (Table 3), Amarasinghe et al. 2022: 417 (Table 3). Chandramouli et al. 2023: 505 (Table 2) compare the species on the Nicobar islands and B. cristatella.

Key to the species of Bronchocela: Amarasinghe et al. 2022: 420. 
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