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Bronchocela rayaensis GRISMER, WOOD, LEE, QUAH, ANUAR, NGADI & SITES, 2015

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Gunung Raya Green-crested Lizard 
SynonymBronchocela rayaensis GRISMER, WOOD, LEE, QUAH, ANUAR, NGADI & SITES 2015
Bronchocela cristatela — ZIMMERER 2004: 87
Bronchocela cristatella — MANTHEY 2008: 49
Bronchocela cristatella — LIM et al. 2009: 23
Bronchocela cristatella — GRISMER 2011: 143
Bronchocela rayaensis — GRISMER & QUAH 2019 
DistributionPeninsular Malaysia (Pulau Langkawi, Tuba Island)

Type locality: Gunung Raya, Pulau Langkawi, Kedah, Peninsular Malaysia (06°22.096 N; 99°49.004 E), 834 m elevation.  
Reproductionoviparous. The paratype (LRCUKM 0136) is a gravid female (SVL 85.0 mm) carrying two eggs. 
TypesHolotype: LSUHC 7535, adult male, collected on 18 August 2005 by L. Lee Grismer and Perry L. Wood Jr., at 2140 hrs Paratype. LRCUKM 0136 (=LSUHC 12061) collected on 2 May 2014 by Ehwan N. at 1000 hrs at 400 m a.s.l. from Gunung Raya, Pulau Langkawi, Kedah, Peninsular Malaysia (06°23.035 N; 99°47.443 E). 
DiagnosisDiagnosis: A small species reaching 85 mm SVL; Tympanum small (DT/DO 0.46–0.49); head squarish in lateral profile (HD/HL 0.49–0.56), relatively narrow (HW/HL 0.56–0.59); forelimbs relatively long (FLL/SVL 0.59); foot relatively short (FoL/HLL 0.37–0.38); hind limbs relatively long (HLL/SVL 0.91–0.99); 5–7 loreal scales; three postmentals; nuchal crest low, longer than diameter of orbit, spines lancolate; 67–71 midbody scales; no rows of paravertebral scales bearing keels pointing dorsoposteriorly; 5–7 rows of dorsal scales bearing keels pointing posteriorly; ventral scales less than five times the width of dorsal scales; 30 subdigital lamellae on the third finger; 31–33 subdigital lamellae on the fourth toe; third finger longer than fourth finger; fourth finger not longer than fifth toe; no white patch between tympanum and orbit; tympanum green in adults; no white or yellow supralabial stripe; no red gular patch; no white ventrolateral line on body; no postfemoral red line; and faint caudal bands. These characters are scored across all species in Tables 4 and 5 in GRISMER et al. 2015. 
Comment 
EtymologyThe specific epithet is derived from the Malay word raya meaning “great” and the Latin suffix-ensis meaning “originating in” and refers to this species discovery at the type locality on Gunung Raya. 
References
  • AMARASINGHE, ; A.A. THASUN; IVAN INEICH, AWAL RIYANTO, JAKOB HALLERMANN, NOVIAR ANDAYANI, A. ABINAWANTO, JATNA SUPRIATNA 2022. Taxonomy and distribution of a common arboreal lizard, Bronchocela jubata Duméril & Bibron, 1837 (Reptilia: Agamidae), with designation of its lectotype from Java, Indonesia. Zootaxa 5150 (1): 065–082 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; P.L. WOOD, JR., CHEOL HAENG LEE, EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & JACK W. SITES, JR. 2015. An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa 3948 (1): 001–023 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., ANCHELEE AOWPHOL, MICHAEL COTA, MATHEW L. MURDOCH, CÉSAR AGUILAR, & MARTA S. GRISMER 2016. Taxonomy, phylogeny, and distribution of Bronchocela rayaensis (Squamata: Agamidae) on the Thai-Malay Peninsula. Zootaxa 4092 (3): 414–420 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Lim, B.L., Noor, A.W.O., Chan, K.O., Belabut, D. & Norhayati, A. 2009. An updated checklist of the herpetofauna of Pulau Singa Besar, Langkawi, Peninsular Malaysia. Malaysian Applied Biology, 39: 13–33 - get paper here
  • Manthey U 2008. Agamid lizards of Southern Asia, Draconinae 1. Terralog 7, 160 pp.
  • QUAH, EVAN S. H.; PERRY L. JR. WOOD, M.S. SHAHRUL ANUAR, MOHD ABDUL MUIN 2020. A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia with an updated checklist of the herpetofauna of Tuba Island. Zootaxa 4767 (1): 138–160 - get paper here
  • Zimmerer, J. 2004. Nature Guide, Langkawi. Sakti Mega Enterprise, Malaysia, 184 pp
 
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