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Calamaria nebulosa LEE, 2021

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Higher TaxaColubridae, Calamariinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Clouded Reed Snake 
SynonymCalamaria nebulosa LEE 2021
Calamaria yunnanensis – STUART & HEATWOLE, 2008
Calamaria yunnanensis – QUAH et al., 2008 (in part)
Calamaria cf. yunnanensis – WEINELL et al. 2021 
DistributionLaos (Phongsaly Province)

Type locality: Laos, Phongsaly Province, Phongsaly District, on the mountainous road to Udomxai [= Oudomxai] about 25 km from Phongsaly City (21º29’N, 102º12’E; ~1000 meters above sea level)  
Reproductionoviparous (presumed, fide Lee, 2021) 
TypesHolotype: FMNH 258666 (field tag: HKV 64382), an adult female collected dead on road (DOR) by Mr. Bounthavy Phommachanh on 6 October 1999. 
DiagnosisDiagnosis: A species of Calamaria distinguished from all other members of the genus from mainland Southeast Asia by having the following combination of morphological characters: (1) rostral wider than high; (2) portion of rostral scale visible from above approximately half the length of the prefrontal suture; (3) six scales and shields surrounding the paraparietal scale; (4) absence of a preocular scale; (5) four supralabials, with the second and third touching the eye; (6) five infralabials, the first pair separating the mental scale from the anterior chin shields; (7) 179 ventrals and three gular scales; (8) 22 paired subcaudals; (9) dorsal color pattern in preservative bluish-gray with five indistinct dark-brown longitudinal stripes; (10) ventral surface of tail in preservative uniform yellow, immaculate; and (11) absence of a distinct white nuchal collar posterior to the head. (Lee, 2021)


Additional details (7055 characters) are available for collaborators and contributors. Please contact us for details. 
CommentSimilar Species: Calamaria yunnanensis CHERNOV, 1962.

Distribution: Presumably has a wider distribution extending across northern Laos, northern Thailand, southern Yunnan Province, China and perhaps also Vietnam.

Conservation: Recommended IUCN status: Data Deficient fide. Lee, 2021. 
References
  • Inger, R. F. & H. MARX 1965. The systematics and evolution of the oriental colubrid snakes of the genus Calamaria. Fieldiana: Zoology 49: 1-304. - get paper here
  • Lee, Justin L. 2021. Description of a new species of Southeast Asian reed snake from northern Laos (Squamata: Colubridae: Genus Calamaria F. Boie, 1827) with a revised diagnosis of Calamaria yunnanensis Chernov, 1962. Journal of Natural History, 55:9-10, 531-560, - get paper here
  • Liang, Y. T., Huang, Z. D., Ding, L., Vogel, G., Ananjeva, N. B., Orlov, N. L., ... & Chen, Z. N. 2024. Revalidated after having been described more than a century ago: Calamaria berezowskii Günther, 1896 (Squamata, Colubridae) from Sichuan, Southwestern China. Zoosystematics and Evolution, 100(3): 897-911 - get paper here
  • Marx, H. & R. F. INGER 1955. Notes on the snakes of the genus Calamaria. Fieldiana: Zoology 37: 167 - 209 - get paper here
  • Quah, Evan S H; Shahrul Anuar, Lee L Grismer, Perry L Wood, Siti Azizah Mohd Nor 2019. Systematics and natural history of mountain reed snakes (genus Macrocalamus; Calamariinae). Zoological Journal of the Linnean Society, 188: 1236 - 1276 - get paper here
  • Stuart, B.L. & Heatwole, H. 2008. Country records of snakes from Laos. Hamadryad 33: 97–106 - get paper here
  • Weinell, J. L., Leviton, A. E., & Brown, R. M. 2021. A new species of reed snake, genus Calamaria (Colubridae: Calamariinae), from Mindoro Island, Philippines. Philippine Journal of Systematic Biology, DOI 10.26757/pjsb2020b14006 - get paper here
 
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